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More reports for this regionA report of mammals and birds recorded
in and around Manu National Park, Peru
August-October 2001
Joe Tobias and Nat Seddon
131 Catharine Street, Cambridge CB1 3AP, UK
email: joetobias22@hotmail.com;
Itinerary
Flight from Cuzco to Boca Manu (BM), boat to Pakitza guard-station (PK): 02/08/01
Boat from Pakitza guard-station to Cocha Cashu Biological Station: 03/08/01
Cocha Cashu Biological Station (CC): 03/08/01 - 09/10/01
Boat to Boca Manu: 09/10/01
Boat from Boca Manu to Pantiacolla lodge: 10/10/01
Pantiacolla lodge (PA): 10/10/01 - 15/10/01
Boat to Manu Road and transport to Cock-of-the-Rock lodge, San Pedro (SP): 15/10/01
Lift to Pillahuarta (PI): 23/10/01
Lift to Cuzco (CU): 24/10/01
Macchu Picchu (MP): 26/10/01
Travel to Ollantaytambo (OL): 27/10/01
Abra Malaga (AM): 28/10/01
Notes on sites
Cocha Cashu Biological Station (c.300 m): we spent several weeks here, focusing on trails near the lake itself, and several times walking out to (and camping at) 'Playa Bonita, a stream some 8 km away along a good trail to the uplands. We spent three morning at the Ccolpa stream, also at the end of a different long trail which leaves the main grid, and four mornings across the Manu river in terra firme forest where there is a trail that you need a boat to reach, plus two mornings around nearby Cocha Totora (reached by trail). We also spent one day at the 'pantanal', a seasonally flooded grassy area reachable by trail. As most of the time was spent following a group of pale-winged trumpeters for a research project, canopy flocks received relatively little attention. It is impossible to visit Cocha Cashu without research permission, although Cocha Salvador is just a few kilometres downriver and can be visited by tourists. With research permission, charges for staying at Cashu are around US$11 per day, depending whether the cook is there.
Boca Manu (c.300 m): we never did any birding around here, just saw a few things around the town.
Pantiacolla (c.400 m): we divided our time between floodplain/bamboo trails (e.g. Tinamou and Capybara trails), the large loop (Monk Saki trail) that runs through terra firme, and the ridge trail (Mirador trail) where we spent a night camping at the first Mirador (c.900 m). We paid US$15 per night on special rate and camping; not sure how much full board is.
San Pedro (c.1,500 m): not really a settlement on the Manu Road, more the name of a bridge beside which stands the Cock-of-the-Rock lodge. We stayed on a fine sheltered camping platform at this lodge, which is much cheaper than the rooms. A short (c.400 m) loop trail descends to the river from the garden of the lodge; this passes through some good forest. Below this lodge (along the road) forest becomes somewhat patchy, punctuated by areas of secondary growth and clearings, some of which contain interesting patches of bamboo. Above the lodge, there is good forest interrupted by landslides all the way, past the Cloud Forest lodge (c.1,650 m), and up towards Pillahuarta. There are two roadside Cock-of-the-Rock leks, one 'owned by the lower lodge and the other 'owned' by the upper lodge. The lower lek (c.150 m uphill from Cock-of-the-Rock lodge) seemed superior in terms of viewing conditions, and is the one referred to in the following accounts. There is an interesting Inca trail starting between the two lodges and heading up to the 'raptor watchpoint' (really just a major bend in the road c.500 m above Cloud Forest lodge) and passing through some good habitat. Easier to do it backwards from the watchpoint down.
Pillahuarta (c.2,900 m): a couple of huts by the upper Manu Road. Good birding above and below these huts, where it is possible to camp on the floor if you speak to the old madman across the road.
Cuzco (c.2,800 m): we visited the famous ruins overlooking the town on two occasions, catching a taxi up there.
Macchu Picchu (c.2,600 m): we took the morning train from Cuzco and the bus up to the ruins and then walked down through the woods along a steep trail through stunted forest, catching the train back a few hours later.
Ollantaytambo/Abra Malaga (2,000-4,000 m): bus from Cuzco, then jumped on a truck at dawn going up the Abra Malaga road (they leave every day). This dropped us off at the pass (taking two hours to get there) and we walked over to the (highly trashed) Polylepis groves, then down halfway to Ollantaytambo again before finally getting a lift to town.
Highlights
In terms of birds, pride of place goes to the tame group of Pale-winged Trumpeters that we studied at CC for two months. Other highlights were Solitary Eagle, Scarlet-shouldered Parakeet, Pheasant Cuckoo, Rufous-vented Ground-cuckoo, White-throated Jacamar, Collared Puffbird, Curl-crested Araçari, Rufous-headed Woodpecker, Marcapata Spinetail, White-browed Tit-spinetail, Peruvian Recurvebill, Bamboo Antshrike, Ihering's Antwren, Yellow-rumped Antwren (Threatened), Manu Antbird, White-lined Antbird, Goeldi's Antbird, Red-and-white Antpitta, Rufous-fronted Antthrush (Threatened), Black-faced Cotinga, White-cheeked Tody-tyrant, Ash-breasted Tit-tyrant (Threatened), Cinnamon-faced Tyrannulet, Dusky-tailed Flatbill, Golden-collared Tanager, Chestnut-breasted Mountain-finch, Black-billed Seed-Finch, Yellow-shouldered Grosbeak, Selva Cacique (Threatened) and Pale-eyed Blackbird. In terms of mammals, the most impressive were Hoffmann's Two-toed Sloth, Giant Otter and ten species of monkeys (best of which was Emperor Tamarin).
Species Lists
A total of c.32 mammal species and 670 bird species was encountered. None of the sightings below were made in Manu National Park proper; almost all are within the buffer zone or research zone. Sightings from around Cuzco, Macchu Picchu, Ollantaytambo and Abra Malaga are outside the park entirely. The list is heavily weighted towards the tropical zone as only a small proportion of time was spent at high altitudes. Nomenclature and systematics follows Emmons and Feer (1997) for mammals and the existing Manu bird list (on CC website; with amendations) for birds.
Key:
Site codes match those given under Itinirary.
[...] = note or uncertain record
(T) = tape-recorded
Mammals
Common Opossum Didelphis marsupialis
Seen on 2-3 occasions at night near camp at CC
Southern Tamandua Tamandua tetradactyla
Seen on three occasions at CC, twice in the grid and once at 'Playa Bonita'
Brown-throated Three-toed Sloth Bradypus tetradactyla
One encountered crossing the stream at 'Playa Bonita', CC, where it was helpless and could be approached to 0 m
Hoffmann's Two-toed Sloth Choloepus hoffmanni
One seen at night at close range near the camp at CC
Pygmy Marmoset Cebuella pygmaea
One group was seen on 2-3 occasions by its roost tree near the camp at CC
Saddleback Tamarin Saguinus fuscicollis
Fairly common at CC, particularly in terra firme (T)
Emperor Tamarin Saguinus imperator
Seen on c.8 occasions at CC, usually in riverine floodplain forest or bamboo-dominated terra firme
Night Monkey Aotus sp. (either Azara's Night Monkey A. azarae, or Black-headed Night Monkey A. nigriceps)
Fairly frequently seen or heard in the grid at CC, on three occasions in daytime when flushed from roosts in vines or bamboo
Dusky Titi Monkey Callicebus moloch brunneus
Fairly common, especially by voice, at CC (T)
Common Squirrel Monkey Saimiri sciureus
Very common at CC (T)
Brown Capuchin Monkey Cebus apella
Common at CC
White-faced Capuchin Monkey Cebus albifrons
Common at CC
Red Howler Monkey Alouatta seniculus
Common, especially by voice, and tame at CC; also seen at PA (T)
Common Woolly Monkey Lagothrix lagothricha
A group was seen from the first Mirador at PA, and another in forest from the Manu Road above SP
White-bellied Spider Monkey Ateles belzebuth
Very common at CC and heard at PA (T)
Short-eared Dog Atelocynus microtis
One set of very fresh tracks was seen along the stream at 'Playa Bonita', CC
Crab-eating Raccoon Procyon cancrivorus
Never seen, but tracks were found regularly at CC
Kinkajou Potus flavus
c.5 seen at night in the grid at CC
Tayra Eira barbara
Three individuals were watched at close range as they chased each other in the grid at CC
Giant Otter Pteronura brasiliensis
A group of nine spent a week or two on the lake at CC; also seen at Cocha Totora
Ocelot Leopardus pardalis and Jaguar Panthera onca tracks were seen regularly at CC, and Puma Felis concolor tracks were seen on the road above PI, but no cats were seen during the entire time
Brazilian Tapir Tapirus terrestris
Many tracks seen at CC (where heard occasionally crashing through undergrowth at night), and one set of tracks seen at PA. Never seen.
Collared Peccary Tayassu tajacu
Small groups of 1-3 were quite commonly encountered in the grid at CC
White-lipped Peccary Tayassu pecari
Large herds (containing up to 80 individuals) were regularly encountered in the grid at CC
Red Brocket Deer Mazama americana
Regularly seen in the grid at CC, where tame
Southern Amazon Red Squirrel Sciurus spadiceus
Common at CC and PA (T)
Bolivian Squirrel Sciurus ignitus
2-3 seen at CC. One small Sciurus seen in lower-stratum at PA looked more like sanborni, but this identification is left provisional.
Capybara Hydrochaeris hydrochaeris
A few family groups seen on sand-banks along the Manu river
Paca Agouti paca
Regularly seen at night at CC
Brown Agouti Dasyprocta variegata
Fairly common in the grid at CC
Green Acouchy Myoprocta pratti
Individuals regularly encountered scampering about in the leaf-litter at CC
Vizcacha Vizcacha vizcacha
A pair at close range on cliffs next to the Polylepis grove at AM
Brazilian Rabbit Sylvilagus brasiliensis
One seen at night in the grid at CC
Birds
Grey Tinamou Tinamus tao
One flushed at close range at CC, where only heard twice
Black Tinamou Tinamus osgoodi
One (maybe two) heard from the Mirador at dawn, PA (T)
Great Tinamou Tinamus major
Commonly heard and seen at CC (T)
White-throated Tinamou Tinamus guttatus
Heard once or twice at CC
Hooded Tinamou Nothocercus nigrocapillus
1-2 heard around 1,700 m (above SP), but couldn't be enticed into view on steep slopes (T)
Cinereous Tinamou Crypturellus cinereus
Fairly commonly heard at CC, almost always in terra firme (T)
Little Tinamou Crypturellus soui
Seen once in várzea and heard occasionally, usually in terra firme at CC (T)
Brown Tinamou Crypturellus obsoletus
c.8 heard near the first Mirador at PA, many more seen and heard around SP (T)
Undulated Tinamou Crypturellus undulatus
Very common at CC throughout floodplain and várzea forest; also PA (T)
Black-capped Tinamou Crypturellus atrocapillus
Fairly common and responsive to playback along the stream at 'Playa Bonita', CC. Also heard by the Madre de Dios river at PA (T)
Variegated Tinamou Crypturellus variegatus
Only heard twice in the uplands near 'Playa Bonita', CC
Bartlett's Tinamou Crypturellus bartletti
Heard commonly and seen regularly in floodplain forest (very rarely in terra firme) at CC (T)
Least Grebe Tachybaptus dominicus
Small numbers on the lake at CC
Neotropic Cormorant Phalacrocorax brasilianus
Small numbers regular around the lake at CC; also along the Manu and Madre de Dios rivers
Anhinga Anhinga anhinga
Small numbers regular at the lake at CC; also along the Manu and Madre de Dios rivers
Horned Screamer Anhima cornuta
Regular and noisy on sand-banks along the Manu river, and at Cocha Totora, CC (T)
Orinoco Goose Neochen jubata
Regular on sand-banks along the Manu river
Muscovy Duck Cairina moschata
Small numbers regular on the lake at CC; also on sand-banks along the Manu river
Torrent Duck Merganetta armata
Only seen along the river near Macchu Picchu, where this species seemed common
Snowy Egret Egretta thula
Small numbers along Madre de Dios river
Capped Heron Pilherodius pileatus
Regular around the lake at CC, and along rivers
Cocoi Heron Ardea cocoi
Regular around the lake at CC, and fairly common along all rivers
Great Egret Casmerodius albus
Regular around the lake at CC, and along rivers
Cattle Egret Bubulcus ibis
One flock seen in flight near Boca Manu
Striated Heron Butorides striatus
Fairly common around the lake at CC, and along rivers
Agami Heron Agamia agami
Seen c.15 times around the lake at CC, and along small streams within forest nearby
Boat-billed Heron Cochlearius cochlearius
Small numbers seen at night around the lake at CC, and along the stream at 'Playa Bonita'
Fasciated Tiger-Heron Tigrisoma fasciatum
c.10 seen while travelling along the Madre de Dios river in the PA area
Rufescent Tiger-Heron Tigrisoma lineatum
Fairly common around the lake at CC
Green Ibis Mesembrinibis cayennensis
Small numbers regular around the lake at CC
Roseate Spoonbill Ajaia ajaja
One flock of c.15 on a sand-bank downstream of PA on the Madre de Dios river
Wood Stork Mycteria americana
Small groups seen occasionally along the Manu river
Black Vulture Coragyps atratus
Small numbers seen throughout
Turkey Vulture Cathartes aura
Recorded from the lower Manu Road, up to above SP and in the CU area
Greater Yellow-headed Vulture Cathartes melambrotus
Fairly common around CC
King Vulture Sarcoramphus papa
Small numbers recorded at CC, mostly at 'Playa Bonita'
Osprey Pandion haliaetus
Occasional at the lake at CC and along major rivers to PA
Grey-headed Kite Leptodon cayanensis
One watched gliding over the Manu River at CC
Swallow-tailed Kite Elanoides forficatus
Regular in small numbers at CC, PA and SP
Snail Kite Rostrhamus sociabilis
Small numbers occasional at CC and nearby Cocha Totora
Double-toothed Kite Harpagus bidentatus
Regularly seen perched or circling at CC and PA; also one at c.1200 m, below SP
Plumbeous Kite Ictinia plumbea
Regular at CC, PA and below SP
Tiny Hawk Accipiter superciliosus
One watched flying across the Manu river at CC
Plain-breasted Hawk Accipiter ventralis
One seen above PI
Slate-colored Hawk Leucopternis schistacea
Fairly common (at least by voice) in várzea, floodplain and swampy areas near CC (T)
White-browed Hawk Leucopternis kuhli
Seen twice in terra firme across the Manu river from CC, and once near 'Playa Bonita'
Great Black-Hawk Buteogallus urubitinga
Small numbers seen along the Manu river
Black-collared Hawk Busarellus nigricollis
Fairly common around CC and nearby Totora
Solitary Eagle Harpyhaliaetus solitarius
One seen circling over hillside opposite SP
Roadside Hawk Buteo magnirostris
Fairly common throughout lowlands and up to SP (T)
Short-tailed Hawk Buteo brachyurus
One seen flying over the confluence of the Manu and Madre de Dios rivers at BM
White-throated Hawk Buteo albigula
At least two above PI
Variable Hawk Buteo polyosoma
A few around AM
Black-and-white Hawk-Eagle Spizastur melanoleucus
One seen soaring over forested hillsides at c.1,300 m on the Manu Road
Ornate Hawk-Eagle Spizaetus ornatus
Fairly common at CC where two active nests were found (T)
Black Caracara Daptrius ater
Fairly common at CC and PA (T)
Red-throated Caracara Daptrius americanus
Common and very noisy around CC and PA (T)
Laughing Falcon Herpetotheres cachinnans
Regularly heard at dawn and dusk at CC, but never seen (T)
Barred Forest-Falcon Micrastur ruficollis
Regularly heard at CC and seen on c.5 occasions; also heard near the first Mirador at PA (T)
Lined Forest-Falcon Micrastur gilvicollis
Heard on c.3 occasions around CC, and played back into view once (T)
Collared Forest-Falcon Micrastur semitorquatus
Seen on c.5 occasions around CC (possible that one of these was buckleyi, but views were inadequate)
American Kestrel Falco sparverius
Seen above PI and around CU
Bat Falcon Falco rufigularis
Fairly common around the lakes and rivers at CC (T)
Speckled Chachalaca Ortalis guttata
Regular in lakeside or riverside growth at CC and PA (T)
Andean Guan Penelope montagnii
Small numbers seen above SP
Spix's Guan Penelope jacquacu
Fairly common around CC (T)
Blue-throated Piping-Guan Pipile cumanensis
Fairly common around CC and PA (T)
Razor-billed Curassow Mitu tuberosa
Fairly common around CC: seen or heard most days (T)
Marbled Wood-Quail Odontophorus gujanensis
Heard on 2-3 occasions in riverine growth along the Manu river, but never seen
Starred Wood-Quail Odontophorus stellatus
Heard regularly around the grid at CC, and on the path to the uplands, and seen on c.5 occasions (T)
Rufous-sided Crake Laterallus melanophaius
Fairly common around the lakes at Cashu and Totora, where vocal and easily seen at dawn or dusk coming out onto floating vegetation (T)
Grey-breasted Crake Laterallus exilis
Heard infrequently at CC and Totora and never seen
Grey-necked Wood-Rail Aramides cajanea
Fairly common at the fringes of CC (T)
Blackish Rail Pardirallus nigricans
One from a bus near OL
Purple Gallinule Porphyrio martinicus
A feww around CC
Common Moorhen Gallinula chloropus
A few around CC
Sungrebe Heliornis fulica
Regular around the shores of CC and once along the stream at 'Playa Bonita'
Sunbittern Eurypyga helias
Regular around the lake at CC, along the stream at 'Playa Bonita' and one seen on the Manu Road where a bridge crosses a river at c.1,250 m (T)
Limpkin Aramus guarauna
Regular around the lake at CC
Pale-winged Trumpeter Psophia leucoptera
Regularly seen around CC, where there is one very tame group, and alarm calls heard once at PA (T)
Wattled Jacana Jacana jacana
Fairly common at CC, common at Cocha Totora (T)
Greater Yellowlegs Tringa melanoleuca
Two seen at the mouth of the Ccolpa stream, and small numbers elsewhere along the Manu and Madre de Dios rivers
Lesser Yellowlegs Tringa flavipes
A few (c.5) seen at the mouth of the Ccolpa stream, CC
Solitary Sandpiper Tringa solitaria
Small numbers along the Manu river, and the Ccolpa and 'Playa Bonita' streams, CC
Spotted Sandpiper Tringa macularia
Fairly common along major rivers from CC to PA
Pectoral Sandpiper Calidris melanotos
Small numbers seen at the mouth of the Ccolpa stream and on sand-banks along the Manu river
Buff-breasted Sandpiper Tryngites subruficollis
One seen on a grassy sandbank along the Manu river
White-backed Stilt Himantopus melanurus
One small flock downstream of PA on the Madre de Dios river
Collared Plover Charadrius collaris
Fairly common on sand-banks along the Manu river
Pied Lapwing Vanellus cayanus
Fairly common along the Manu river
Andean Lapwing Vanellus resplendens
2-3 in the open valley floor at AM
Andean Gull Larus serranus
One seen flying over fields near the pass at AM
Large-billed Phaetusa simplex
Large numbers breeding on sand-banks along the Manu river
Yellow-billed Tern Sterna superciliaris
Small numbers breeding on sand-banks along the Manu river
Black Skimmer Rynchops niger
Several fairly large colonies along the Manu river
Spot-winged Pigeon Columba maculosa
A few around CU
Band-tailed Pigeon Columba fasciata
Regular in cloud forest above SP and around PI
Pale-vented Pigeon Columba cayennensis
Common in lakeside or riverside forest at CC and PA (T)
Plumbeous Pigeon Columba plumbea
Common at CC, PA and right up to nearly 2,000 m on Manu Road (T)
Ruddy Pigeon Columba subvinacea
Common at CC and PA (T)
Eared Dove Zenaida auriculata
Common around CU and OL
Blue Ground-Dove Claravis pretiosa
Occasional at PK and CC (T)
Grey-fronted Dove Leptotila rufaxilla
Common, particularly in waterside forest, at CC and PA (T)
Sapphire Quail-Dove Geotrygon saphirina
One seen walking on forest floor a short way up the Mirador trail at PA
White-throated Quail-Dove Geotrygon frenata
Several calling around PI, others heard at SP were probably this species (T)
Ruddy Quail-Dove Geotrygon montana
Fairly common at CC and PA (T)
Blue-and-yellow Macaw Ara ararauna
Common at CC and PA (T)
Scarlet Macaw Ara macao
Regular at CC, several pairs nesting in palms in swampy areas (T)
Red-and-green Macaw Ara chloropterus
Regular at CC and PA
Chestnut-fronted Macaw Ara severa
Fairly common at CC and PA (T)
Blue-headed Macaw Ara couloni
Never seen well, but it's double note was heard on a few occasions over forest at CC
White-eyed Parakeet Aratinga leucophthalmus
A few at CC and PA
Dusky-headed Parakeet Aratinga weddellii
Fairly common at CC and PA (T)
Mitred Parakeet Aratinga mitrata
Only identified below Macchu Picchu where it seemed common
Painted Parakeet Pyrrhura picta
Common at CC and PA; never saw rupicola and this was definitely in the minority (T)
Andean Parakeet Bolborhynchus orbygnesius
One flock of c.25 at AM
Dusky-billed Parrotlet Forpus sclateri
Fairly common along the streams near the Ccolpa and at 'Playa Bonita' where they visit clay-licks; never seen in the forest itself (T)
Cobalt-winged Parakeet Brotogeris cyanoptera
Very common throughout the forest at CC and PA (T)
Tui Parakeet Brotogeris sanctithomae
Regular around the lakeshore and riverside at CC (T)
Scarlet-shouldered Parrotlet Touit huetii
One pair seen at PA where digging a cavity into an arboreal termitary (T)
White-bellied Parrot Pionites leucogaster
Common and vocal at CC and PA (T)
Blue-headed Parrot Pionus menstruus
Fairly common and vocal at CC and PA (T)
Yellow-crowned Parrot Amazona ochrocephala
Regular around CC (T)
Scaly-naped Parrot Amazona mercenaria
Flocks flying at dusk over SP daily
Mealy Parrot Amazona farinosa
Common and noisy at CC and PA (T)
Yellow-billed Cuckoo Coccyzus americanus
Two seen with canopy flocks towards the end of our stay at CC
Dark-billed Cuckoo Coccyzus melacoryphus
One seen in lakeside trees at CC, another seen in floodplain forest at PA
Squirrel Cuckoo Piaya cayana
Common in lower-lying areas (T)
Hoatzin Opisthocomus hoazin
Common around CC and Totora (T)
Greater Ani Crotophaga major
Fairly common around CC and Totora
Smooth-billed Ani Crotophaga ani
Seen around PA and the lower Manu Road
Pheasant Cuckoo Dromococcyx phasianellus
Regularly heard in bamboo along the stream at 'Playa Bonita', where seen once very well in response to playback (T)
Pavonine Cuckoo Dromococcyx pavoninus
Heard twice in bamboo in terra firme across the river from CC (T)
Rufous-vented Ground-Cuckoo Neomorphus geoffroyi
Prolonged views at close range by the station at CC, and at the beginning of the Tinamou trail at PA
Barn Owl Tyto alba
One seen in flight shortly after dusk at PI
Tropical Screech-Owl Otus choliba
Heard twice at the lake at CC
Rufescent Screech-Owl Otus ingens
Heard regularly around SP and responded strongly to playback in the grounds of the lodge (T)
Tawny-bellied Screech-Owl Otus watsonii
Heard commonly at CC and PA; seen once each at both these sites (T)
Vermiculated Screech-Owl Otus vermiculatus
2-3 heard near the top of the Mirador trail, PA (T)
Mottled Owl Strix virgata
Heard occasionally in forest at CC and once at PA (T)
Black-banded Owl Strix huhula
Heard once at CC
Crested Owl Lophostrix cristata
Heard nightly at CC and PA; seen three times at close range in daylight at CC (T)
Spectacled Owl Pulsatrix perspicillata
Heard often at CC, and seen once in daylight (T)
Band-bellied Owl Pulsatrix melanota
Two pairs heard near the first Mirador at PA; came very close to playback but couldn't be seen (T)
Amazonian Pygmy-Owl Glaucidium hardyi
Commonly heard and regularly seen in daylight at CC (T)
Ferruginous Pygmy-Owl Glaucidium brasilianum
Heard occasionally at CC, mostly at 'Playa Bonita' (T)
Great Potoo Nyctibius grandis
Heard occasionally at CC but never seen (T)
Long-tailed Potoo Nyctibius aethereus
Heard on c.5 occasions at CC but never seen; one found incubating eggs at PA (T)
Grey Potoo Nyctibius griseus
Heard twice at CC in riverine Cecropias
Short-tailed Nighthawk Lurocalis semitorquatus
Seen at dusk from a canoe flying over forest at CC
Rufous-bellied Nighthawk Lurocalis rufiventris
One bird played in to close range at dusk at PI (T)
Sand-colored Nighthawk Chordeiles rupestris
Common on sand-banks along the Manu river, and several visited the lake itself at CC each dusk
Pauraque Nyctidromus albicollis
Fairly common around the lake at CC, and even quite deep in the forest (T0
Ocellated Poorwill Nyctiphrynus ocellatus
Common in forest at CC: often heard and c.10 nests found by eyeshine at night (T)
Lyre-tailed Nightjar Uropsalis lyra
2-3 heard at c.1,700 m above SP; two males seen well and one female found in daylight on a roadside nest (T)
Chestnut-collared Swift Streptoprocne rutila
Common in large flocks at SP
White-collared Swift Streptoprocne zonaris
Occasional over CC and fairly common along the Manu Road
Grey-rumped Swift Chaetura cinereiventris
Fairly common at CC
Pale-rumped Swift Chaetura egregia
Regular at CC, in small numbers over the 'Playa Bonita' stream, and seen once at PA
Short-tailed Swift Chaetura brachyura
Fairly common at CC
Fork-tailed Palm-Swift Tachornis squamata
Small numbers seen at CC
Rufous-breasted Hermit Glaucis hirsuta
Common at CC where several nests were found
Pale-tailed Barbthroat Threnetes leucurus
A few seen at CC and several seen at PA (T)
Green Hermit Phaethornis guy
2-3 seen at SP
Great-billed Hermit Phaethornis malaris
Seen rarely, or at least identified rarely in terra firme at CC
White-bearded Hermit Phaethornis hispidus
Common and vocal at CC (T)
Koepcke's Hermit Phaethornis koepckeae
Several seen along the Mirador trail at PA
Needle-billed Hermit Phaethornis philippii
One seen in bamboo near the river on the Tinamou trail at PA
Reddish Hermit Phaethornis ruber
Common at CC, PA and seen below SP (T)
[White-browed Hermit Phaethornis stuarti]
Possibly seen on several occasions but we found this species impossible to identify with certainty
Buff-tailed Sicklebill Eutoxeres condamini
Seen once below SP
Green-fronted Lancebill Doryfera ludovicae
Seen regularly at flowering trees near SP
Grey-breasted Sabrewing Campylopterus largipennis
Seen on 2-3 occasions at CC
White-necked Jacobin Florisuga mellivora
Seen occasionally at CC (mostly feeding high over the lake or streams), also in the clearing at PA
Green Violet-ear Colibri thalassinus
Small numbers seen between SP and PI
Sparkling Violet-ear Colibri coruscans
Fairly common between SP and PI (T)
Blue-tailed Emerald Chlorostilbon mellisugus
A female seen well at CC in August
Fork-tailed Woodnymph Thalurania furcata
Fairly common at CC and PA
White-chinned Sapphire Hylocharis cyanus
Fairly common in more open forest or clearings at CC (T)
Golden-tailed Sapphire Chrysuronia oenone
Seen once at SP
Many-spotted Hummingbird Taphrospilus hypostictus
Seen once at SP in roadside low canopies, fighting gamely with a Sparkling Violetear
Green-and-white Hummingbird Amazilia viridicauda
Singing around the ruins at MP
Sapphire-spangled Emerald Amazilia lactea
A few seen and heard along the lower Manu Road (T)
Speckled Hummingbird Adelomyia melanogenys
Common and vocal in forest around SP (T)
Gould's Jewelfront Heliodoxa aurescens
Seen on c.5 occasions at CC
Violet-fronted Brilliant Heliodoxa leadbeateri
Fairly common at SP, visiting feeders at Cock-of-the-rock lodge
Giant Hummingbird Patagona gigas
1-2 seen at AM
Shining Sunbeam Aglaeactis cupripennis
Fairly common at AM
White-tufted Sunbeam Aglaeactis castelnaudi
c.4 seen in shrubbery by the road between OL and AM
Great Saphirewing Pterophanes cyanopterus
c.15 seen below AM
Violet-throated Starfrontlet Coeligena violifer
1 seen at c.2,000 m (above SP), and another 3-4 seen at c.3,100 m (above PI)
Sword-billed Hummingbird Ensifera ensifera
One seen well above PI
Chestnut-breasted Coronet Boissonneaua matthewsii
One seen in woodland below MP
Amethyst-throated Sunangel Heliangelus amethysticollis
Fairly common in woodland near PI
Coppery-naped Puffleg Eriocnemis sapphiropygia
One seen at c.2,000 m (above SP)
Booted Racquet-tail Ocreatus underwoodii
Small numbers seen in roadside vegetation above SP
Green-tailed Trainbearer Lesbia nuna
A few seen near CU
Scaled Metaltail Metallura aeneocauda
One seen well at PI
Tyrian Metaltail Metallura tyrianthina
Race with blue (not reddish) tail common above c.1,800 m on Manu Road
Blue-mantled Thornbill Chalcostigma stanleyi
c.5 seen around Polylepis at AM
Long-tailed Sylph Aglaiocercus kingi
Fairly common above SP
Wedge-billed Hummingbird Schistes geoffroyi
One seen in song c.300 m downhill from Cloud Forest lodge, Manu Road (T)
Black-eared Fairy Heliothryx aurita
2-3 seen in canopies at CC
Long-billed Starthroat Heliomaster longirostris
One at CC (edge of Cocha Totora)
White-bellied Woodstar Acestrura mulsant
One male at SP
Crested Quetzal Pharomachrus antisianus
2-3 between SP and Cloud Forest lodge (T)
Golden-headed Quetzal Pharomachrus auriceps
c.8 seen above SP and below PI; quite often heard (T)
Black-tailed Trogon Trogon melanurus
Very common at CC, less so at PA (T)
White-tailed Trogon Trogon viridis
Scarce at CC, more common at PA (T)
Collared Trogon Trogon collaris
Fairly common at CC and PA
Masked Trogon Trogon personatus
One in song above PI (T)
Blue-crowned Trogon Trogon curucui
Fairly common at CC and a few at PA (T)
Violaceous Trogon Trogon violaceus
Scarce at CC, more common at PA and lower Manu Road (T)
Ringed Kingfisher Ceryle torquata
Common at lakes and rivers in the lowlands
Amazon Kingfisher Chloroceryle amazona
A few around CC and along the Manu and Madre de Dios rivers
Green Kingfisher Chloroceryle americana
Fairly common at CC, including along small forest streams
Green-and-rufous Kingfisher Chloroceryle inda
Seen occasionally at CC
American Pygmy Kingfisher Chloroceryle aenea
Seen occasionally at CC
Broad-billed Motmot Electron platyrhynchum
Seen c.5 times and heard fairly often at CC (T)
[Rufous Motmot Baryphthengus martii]
Probably heard at PA, but never definitely recorded
Blue-crowned Motmot Momotus momota
Common at CC and PA (T)
Highland Motmot Momotus aequatorialis
Seen on c.4 occasions at SP
Chestnut Jacamar Galbalcyrhynchus purusianus
Heard regularly and seen on c.6 occasions in swampy or lakeside forest at CC
White-throated Jacamar Brachygalba albogularis
Fairly common along the Ccolpa and 'Playa Bonita' streams, but not seen anywhere else at CC (T)
Bluish-fronted Jacamar Galbula cyanescens
Common, especially in streamside, riverside or lakeside vegetation at CC; seen up to around 1,400 m on the Manu road (T)
Great Jacamar Jacamerops aureus
Heard regularly and seen occasionally at CC (T)
White-necked Puffbird Notharchus macrorhynchos
Never seen, but occasionally heard calling from upper canopy at CC (T)
Chestnut-capped Puffbird Bucco macrodactylus
Only seen once, at a clearing in the grid at CC
Collared Puffbird Bucco capensis
Seen on c.5 occasions in the grid at CC, usually at close range in lower-stratum; heard only once, after dusk (T)
Striolated Puffbird Nystalus striolatus
Heard regularly at CC and seen on c.5 occasions, usually in mid-levels of vine-tangled forest near the Manu river
Semicollared Puffbird Malacoptila semicincta
Seen on c.8 occasions in lower-stratum at CC
Rufous-capped Nunlet Nonnula ruficapilla
Seen regularly in riverine regrowth along the Manu river and smaller streams (e.g. at 'Playa Bonita' and Ccolpa)
Black-fronted Nunbird Monasa nigrifrons
Very common in floodplain forest at CC and PA (T)
White-fronted Nunbird Monasa morphoeus
Scarce in floodplain forest and slightly more common in terra firme at CC and PA (T)
Swallow-winged Puffbird Chelidoptera tenebrosa
Common in riverside vegetation throughout the lowlands; many nests found along the 'Playa Bonita' and Ccolpa streams (T)
Gilded Barbet Capito auratus
Very common by voice in canopies at CC and PA, much less often seen (T)
Lemon-throated Barbet Eubucco richardsoni
Fairly common by voice at CC and PA, seen regularly
Scarlet-hooded Barbet Eubucco tucinkae
1-2 seen along the 'Playa Bonita' stream, and several in riverside Cecropias at PA
Versicolored Barbet Eubucco versicolor
2-3 seen above SP, around 2,200 m
Emerald Toucanet Aulacorhynchus prasinus
A few at CC, mostly around the 'Playa Bonita' and Ccolpa streams
Chestnut-tipped Toucanet Aulacorhynchus derbianus
Not seen, but one taped while calling on the Manu Road at c.1,200 m
Blue-banded Toucanet Aulacorhynchus coeruleicinctis
c.8 seen above SP, between 1,750 and 2,400 m
Lettered Aracari Pteroglossus inscriptus
Fairly common at CC and PA
Brown-mandibled Aracari Pteroglossus mariae
Fairly common at CC
Chestnut-eared Aracari Pteroglossus castanotis
Common at CC and PA (T)
Curl-crested Araçari Pteroglossus beauharnaesii
Seen on c.15 occasions at CC
Golden-collared Toucanet Selenidera reinwardtii
Fairly common by voice at CC, but only seen on c.10 occasions
Channel-billed Toucan Ramphastos vitellinus
Fairly common by voice at CC (T)
White-throated Toucan Ramphastos tucanus
Very common at CC and fairly common at PA (T)
Rufous-breasted Piculet Picumnus rufiventris
Seen or heard on c.10 occasions in swampy or bamboo-dominated forest at CC (T)
Yellow-tufted Woodpecker Melanerpes cruentatus
Common at CC and PA (T)
Little Woodpecker Veniliornis passerinus
Fairly common at CC, PA, and the lower Manu Road, mostly in riverside habitat (T)
Red-stained Woodpecker Veniliornis affinis
Fairly common with mid-stratum flocks at CC
[White-throated Woodpecker Piculus leucolaemus]
Possibly seen at CC, but all good views were of the next species
Golden-green Woodpecker Piculus chrysochloros
Often seen and heard at CC (T)
Golden-olive Woodpecker Piculus rubiginosus
Seen on c.8 occasions above SP
Spot-breasted Woodpecker Colaptes punctigula
Seen or heard on c.4 occasions in riverine or lakeside habitat at CC (T)
Andean Flicker Colaptes rupicola
Fairly common at AM
Scaly-breasted Woodpecker Celeus grammicus
Regularly seen, and heard almost daily at CC (T)
Chestnut Woodpecker Celeus elegans
Seen on c.15 occasions at CC, and once at PA (T)
Cream-colored Woodpecker Celeus flavus
Seen or heard on c.10 occasions at CC, always around swampy areas of floodplain forest in grid (T)
Rufous-headed Woodpecker Celeus spectabilis
Heard on most visits to the 'Playa Bonita' stream, where it was seen on 3 occasions in streamside cane and bamboo (T)
Lineated Woodpecker Dryocopus lineatus
Occasionally heard in flooded areas with dead trees at CC, also at PA and lower Manu Road
Red-necked Woodpecker Campephilus rubricollis
Fairly common at CC and PA (T)
Crimson-crested Woodpecker Campephilus melanoleucos
Fairly common - but less so than the last species - at CC
Plain-brown Woodcreeper Dendrocincla fuliginosa
Fairly common, at least by voice, at CC and PA (T)
White-chinned Woodcreeper Dendrocincla merula
Seen or heard (a quiet tittering) on c.20 occasions at CC - accompanying most ant-swarms (T)
Long-tailed Woodcreeper Deconychura longicauda
1-2 heard most days in the grid at CC (T)
Olivaceous Woodcreeper Sittasomus griseicapillus
Regularly seen and heard accompanying mid-stratum mixed-species flocks at CC and PA (T)
Wedge-billed Woodcreeper Glyphorynchus spirurus
Fairly common at CC, but infrequently heard
Long-billed Woodcreeper Nasica longirostris
Regular around the lake at CC (T)
Cinnamon-throated Woodcreeper Dendrexetastes rufigula
Regularly heard at CC and PA, but only seen on c.5 occasions at the former (T)
Strong-billed Woodcreeper Xiphocolaptes promeropirhynchus
Lawrence's Thrushes Turdus lawrencii were regularly heard imitating this species at CC, but the real thing was only heard twice
Barred Woodcreeper Dendrocolaptes certhia
Fairly common at CC, especially by voice (T)
Black-banded Woodcreeper Dendrocolaptes picumnus
Appeared much less common than the previous species, only seen or heard on c.3 occasions at CC
Straight-billed Woodcreeper Xiphorhynchus picus
Common in lakeside or riverside habitat at CC and PA (T)
Striped Woodcreeper Xiphorhynchus obsoletus
Heard regularly in swampy woodland with stands of Heliconia, or in lakeside vegetation at CC (T)
Tschudi's Woodcreeper Xiphorhynchus chunchotambo
Definitely saw this twice at CC, and it was probably heard occasionally
Elegant Woodcreeper Xiphorhynchus elegans
Commonly heard and regularly seen with lower-stratum flocks at CC and PA (T)
Buff-throated Woodcreeper Xiphorhynchus guttatus
Very common and vocal at CC and PA (T)
Olive-backed Woodcreeper Xiphorhynchus triangularis
Seen on c.3 occasions with mixed-species flocks above SP
Lineated Woodcreeper Lepidocolaptes albolineatus
Seen regularly accompanying canopy flocks at CC and PA, heard infrequently (T)
Red-billed Scythebill Campylorhamphus trochilirostris
Seen in bamboo-dominated streamside woodland at 'Playa Bonita' and in taller terra firme at PA (T)
Bar-winged Cinclodes Cinclodes fuscus
Fairly common at AM
White-winged Cinclodes Cinclodes atacamensis
2-3 at AM
Pale-legged Hornero Furnarius leucopus
Fairly common at CC and PA in lakeside, riverside or streamside habitats (T)
full species list continues on next page - click here