Peru - The Manu Biosphere Reserve, April 2019

Published by Manu Expeditions (birding AT manuexpeditions.com)

Participants: Gustavo Bautista

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ITINERARY:

April 8th: We started early after breakfast, leaving from Cusco to spend the whole day birding around the high Andean lake called Huacarpay. Overnight in Cusco.

April 9th to 10th: Early departure to go to Manu. First birding stop was near Huancarani where we looked for a few endemics like Bearded Mountaineer and Creamy-crested Spinetail, before driving to the picturesque town of Paucaro. From there it was one hour going up to get to the entrance of the Manu National Park where we had our picnic lunch, afterwards we started birding all the way down to Wayqecha Lodge where we spent the night. The next day after an early hearty breakfast we birded up and down the road looking for species we may have missed the day before, and also spent time watching hummingbirds near the lodge. Both nights at Wayqecha Lodge.

April 11th to 14th: Leaving after breakfast from Wayqecha we travelled down to Cock-of-the-Rock Lodge. On the way down we made several stops for birding and enjoyed our picnic lunch. We arrived in good time so we could enjoy all the birds that attend the feeding stations and especially the hummingbirds at the feeders. The following days we birded at different elevations up and down the road looking for target birds as well enjoying watching different species of tanagers and hummingbirds right in front of the dinning room. Overnights at Cock-of-the-Rock Lodge.

April 15th: Leaving the lodge after breakfast we continued going down the road to Villa Carmen Biological Station. This lodge has one of the most amazing bamboo thickets, so great to look for bamboo specialists and also there is a really nice garden with feeders, so a place to relax and watch birds and having a drink at the same time. Good owls around the lodge too. Overnight at Villa Carmen Lodge.

April 16th to 18th: We left the lodge after lunch, after birding a bit more in the garden and along the bamboo trails. We continued going down the Manu road, stopping in important areas until we arrived to the river port of Atalaya, where we took our private motorized boat down to Amazonia Lodge which is at ca. 600m. We spent 3 nights here. The main attractions here are the hummingbirds that attend the feeders and the whole garden itself that has got great bird activity most of the day and to visit the trail system of the lodge. Overnights at Amazonia Lodge.

April 19th to 24th: After birding a bit more in the garden after breakfast, we got ready for our 6 to 8 hours (including stops) boat ride birding along the Madre de Dios River to get to Manu Wildlife Center. We arrived in daylight so we birded the grounds of the lodge for a bit. In the following 6 days we would be birding the lodge’s amazing trail system. One day we left the lodge to go to the Blanquillo claylick to watch the macaws, a spectacular event Other days we went with the boat downriver to visit two oxbow lakes, Camungo and Blanco, one of them has a very tall canopy tower which is great for watching mixed species foraging flocks. Nearby the Manu Wildlife Center Lodge there is a Canopy tower as well, which is good to visit at any time.

April 25th: On this day we had a long trip to get from MWC (Manu Wildlife Center) to Puerto Maldonado. We left early in the morning after breakfast to visit the Antthrush Trail to look for more bamboo specialists we may have missed and then continued going down the Madre de Dios River until we arrived at Colorado Port, where we took private cars for an hour's ride towards the Inambari river, then another boat just to go across the river and on the other side of the river our private bus was waiting. It is a long day traveling but birding mode is still ON! Overnight in Cabaña Quinta in Puerto Maldonado.

April 26th: After breakfast we went in our van to bird nearby Puerto Maldonado in 2 specific hotspots: La Pastora road and La Cachuela road. After birding we had time to go back to the hotel to get ready to leave to the airport to catch our flights back to Cusco or to Lima.

SPECIES LIST

TINAMOUS


Great Tinamou (Tinamus major). Heard only.

Cinereous Tinamou (Crypturellus cinereus). Heard only.

Little Tinamou (Crypturellus soui). Seen at Amazonia Lodge.

Brown Tinamou (Crypturellus obsoletus). Heard only.

Undulated Tinamou (Crypturellus undulatus). Heard only.

Black-capped Tinamou (Crypturellus atrocapillus). Heard only.

Bartlett's Tinamou (Crypturellus bartletti). Heard only.

SCREAMERS

Horned Screamer (Anhima cornuta). Heard and seen along the Madre de Dios river and oxbow lakes.

DUCKS, GEESE, AND WATERFOWL

Orinoco Goose (Oressochen jubatus). A pair seen along the Madre de Dios river.

Muscovy Duck (Cairina moschata). Seen along the Madre de Dios river.

Torrent Duck (Merganetta armata). Seen at Manu Cloud forest Lodge.

Puna Teal (Spatula puna). Seen at Huacarpay lakes.

Cinnamon Teal (Spatula cyanoptera). Seen at Huacarpay lakes.

Yellow-billed Pintail (Anas georgica). Seen at Huacarpay lakes.

Yellow-billed Teal (Anas flavirostris). Seen at Huacarpay lakes. The SACC says “Hellmayr & Conover (1948a) and many earlier classifications treated andium (Andean Teal) as a separate species from A. flavirostris. Many authors, from Meyer de Schauensee (1970) to Dickinson (2003), have treated andium as a subspecies of A. flavirostris. Anas andium was considered a separate species from Anas flavirostris by Ridgely et al. (2001), and this was followed by Hilty (2003). SACC proposal passed to recognize andium as separate species. 7 Jaramillo (2003) further suggested that the subspecies oxyptera may also deserve recognition as a separate species from A. flavirostris.

GUANS

Speckled Chachalaca (Ortalis guttata). A very common bird of Amazonian lowlands. Common in more lightly wooded and forest edge areas in the lowlands.

Andean Guan (Penelope montagnii). Seen at Rocotal area (Manu road).

Spix's Guan (Penelope jacquacu). Seen at Manu Wildlife Center. Named after the German naturalist and collector in Brazil (1781-1826) Johann Baptist Ritter von Spix.

Blue-throated Piping-Guan (Pipile cumanensis). Seen at Amazonia Lodge, Manu Wildlife Center and along the Madre de Dios river.

NEW WORLD QUAIL

Rufous-breasted Wood-Quail (Odontophorus speciosus). Heard and seen at Cock-of-the-Rock Lodge.

GREBES

White-tufted Grebe (Rollandia rolland). Seen at Huacarpay lakes. Rollandia: Specific name Podiceps rolland. rolland= In honor of Thomas Pierre Rolland, Master Gunner (1776-1847) French Navy, of the corvette L’Uranie which circled the globe 1817-1848.

CORMORANTS

Neotropic Cormorant (Phalacrocorax brasilianus). A common species seen along the Madre de Dios river.

DARTERS

Anhinga (Anhinga anhinga). Seen at Cocha Camungo and Cocha Blanco (oxbow lakes).

HERONS, EGRETS, AND BITTERNS

Rufescent Tiger-Heron (Tigrisoma lineatum). Seen at Cocha Blanco (oxbow lake).

Fasciated Tiger-Heron (Tigrisoma fasciatum). A fairly common species seen along the Madre de Dios river.

Black-crowned Night-Heron (Nycticorax nycticorax). Seen at Huacarpay lakes.

Cocoi Heron (Ardea cocoi). A common species seen along the Madre de Dios river.

Great Egret (Ardea alba). A common species seen along the Madre de Dios river.

Snowy Egret (Egretta thula). A common species seen along the Madre de Dios river.

Little Blue Heron (Egretta caerulea). A boreal migrant? seen along the Madre de Dios river.

Cattle Egret (Bubulcus ibis). Common species.

Striated Heron (Butorides striata). Seen at Cocha Camungo (oxbow lake).

Capped Heron (Pilherodius pileatus). Seen along the Madre de Dios river.

IBISES

Puna Ibis (Plegadis ridgwayi). Seen near Huacarpay lakes.

Green Ibis (Mesembrinibis cayennensis). Seen at Cocha Camungo.

Roseate Spoonbill (Ajaia ajaja). Seen along the Madre de Dios river.

NEW WORLD VULTURES

Black Vulture (Coragyps atratus). Common species.

Turkey Vulture (Cathartes aura). Common species. Jaramillo (2003) suggested that the resident tropical subspecies ruficollis and the southern subspecies group (jota and "falklandica") might merit recognition as separate species from the northern Cathartes aura group.

Greater Yellow-headed Vulture (Cathartes melambrotus). Common species in the Amazon.

King Vulture (Sarcoramphus papa). Seen at Amazonia Lodge, Blanquillo claylick, Cocha Camungo (oxbow lake), and along the Madre de Dios river.

HAWKS, EAGLES, AND KITES

Crested Eagle (Morphnus guianensis). Seen at Manu Wildlife Center. Mostly more difficult to see than Harpy Eagle. Named for the country of Guiana.

Black Hawk-Eagle (Spizaetus tyrannus). Heard only.

Black-collared Hawk (Busarellus nigricollis). Seen at Cocha Blanco. A lake and riverside specialist.

Plumbeous Kite (Ictinia plumbea). Common. Seen mainly along the Madre de Dios river.

Bicolored Hawk (Accipiter bicolor). Seen along the Madre de Dios river.

Great Black Hawk (Buteogallus urubitinga). Common. Seen mainly along the Madre de Dios river.

Roadside Hawk (Rupornis magnirostris). Common. Mainly along the Madre de Dios river.

Variable Hawk (Geranoetus polyosoma). Seen along the road on the way from Cusco city to Paucartambo. Farquhar (1988) concluded that Buteo poecilochrous (Puna) and B. polyosoma (Red-backed) are conspecific, as they were formerly treated (REF); he was unable to find any way to reliably diagnose the two forms using plumage characters or measurements. Ridgely & Greenfield (2001), Jaramillo (2003), and Schulenberg et al. (2007) followed this treatment and suggested "Variable Hawk" be retained for the composite species. Genetic data (Riesing et al. 2003) are consistent with hypothesis that B. polyosoma and B. poecilochrous are conspecific.

Black-chested Buzzard-Eagle (Geranoetus melanoleucus). Seen at Huacarpay lakes.

Gray-lined Hawk (Buteo nitidus). Seen along the Madre de Dios river.

Broad-winged Hawk (Buteo platypterus). Seen along the Manu road. A boreal migrant.

SUNBITTERN

Sunbittern (Eurypyga helias). Seen at Amazonia lodge, Cocha Blanco (oxbow lake).

RAILS, GALLINULES, AND COOTS

Gray-breasted Crake (Laterallus exilis). Seen at Cocha Camungo.

Uniform Crake (Amaurolimnas concolor). Seen at Amazonia lodge.

Rufous-sided Crake (Laterallus melanophaius). Seen at Cocha Camungo.

Plumbeous Rail (Pardirallus sanguinolentus). Seen at Huacarpay lakes.

Common Gallinule (Gallinula galeata). Seen at Huacarpay lakes. Vocal, plumage, and genetic data suggest that New World populations of widely distributed Old World Common Moorhen Gallinula chloropus should be treated as a separate species Gallinula galeata (Constantine 2006, Groenenberg et al. 2008).

Gray-cowled Wood-Rail (Aramides cajaneus). Heard only.

Slate-colored Coot (Fulica ardesiaca). Seen at Huacarpay lakes. Called "Andean Coot" in Fjeldså & Krabbe (1990), Taylor (1996 and Ridgely et al. (2001) but other authors use Slate-colored Coot.

STILTS

Black-necked Stilt (Himantopus mexicanus). Seen at Huacarpay.

LIMPKIN

Limpkin (Aramus guarauna). Seen at Cocha Blanco (oxbow lake).

PLOVERS AND LAPWINGS

Collared Plover (Charadrius collaris). Seen along the Madre de Dios river.

Pied Lapwing (Vanellus cayanus). Seen along the Madre de Dios river. A very pretty bird!

Southern Lapwing (Vanellus chilensis). Seen along the Madre de Dios river. A recent colonist from Brazil due to deforestation.

Andean Lapwing (Vanellus resplendens). Seen at Huacarpay lakes.

SANDPIPERS AND ALLIES

Spotted Sandpiper (Actitis macularius). Seen along the Madre de Dios river.

Greater Yellowlegs (Tringa melanoleuca). Seen along the Madre de Dios river.

Lesser Yellowlegs (Tringa flavipes). Seen along the Madre de Dios river.

JACANA

Wattled Jacana (Jacana jacana). Seen at Cocha Camungo and Cocha Blanco (oxbow lakes).

GULLS, TERNS, AND SKIMMERS

Andean Gull (Chroicocephalus serranus). Seen at Huacarpay lakes.

Laughing Gull (Leucophaeus atricilla). Seen at Huacarpay lakes. A vagrant here.

Yellow-billed Tern (Sternula superciliaris). Seen along the Madre de Dios river.

Large-billed Tern (Phaetusa simplex). Seen along the Madre de Dios river.

Black Skimmer (Rynchops niger). Seen along the Madre de Dios river.

PIGEONS AND DOVES

Rock Pigeon (Columba livia). Introduced species and common. Feral population.

Pale-vented Pigeon (Patagioenas cayennensis). Seen at Cocha Blanco (oxbow lake), Puerto Maldonado city and along the Madre de Dios river.

Spot-winged Pigeon (Patagioenas maculosa). Seen at Huacarpay lakes.

Band-tailed Pigeon (Patagioenas fasciata). Fairly common.

Plumbeous Pigeon (Patagioenas plumbea). A common species from the foothills down to the Amazon.

Ruddy Pigeon (Patagioenas subvinacea). Seen at Amazonia Lodge and Manu Wildlife Center. VULNERABLE.

Ruddy Ground-Dove (Columbina talpacoti). Common. Found in open and semi-open areas in the Amazon.

Bare-faced Ground-Dove (Metriopelia ceciliae). Seen at Huacarpay lakes.

Black-winged Ground-Dove (Metriopelia melanoptera). Seen at Huacarpay lakes.

Ruddy Quail-Dove (Geotrygon montana). Seen at Manu Wildlife Center.

Gray-fronted Dove (Leptotila rufaxilla). Common. Seen at Amazonia Lodge and Manu Wildlife Center and near Cocha Camungo (oxbow lake).

Eared Dove (Zenaida auriculata). Common in the Andes.

HOATZIN

Hoatzin (Opisthocomus hoazin). Common in the Amazon. Seen mainly at the oxbow lakes.

CUCKOOS

Smooth-billed Ani (Crotophaga ani). Common in the Amazon. Seen at Patria (600-900m) (Manu road), Blanquillo claylick, Cocha Camungo (oxbow lake), Puerto Maldonado city and along the Madre de Dios river.

Greater Ani (Crotophaga major). Common in the Amazon. Especially along oxbow lakes.

Pavonine Cuckoo (Dromococcyx pavoninus). Seen at Manu Wildlife Center.

Rufous-vented Ground-Cuckoo (Neomorphus geoffroyi). Seen at Villa Carmen. Named for or by Perroquet Geoffroy of Levaillant 1805.

Little Cuckoo (Coccycua minuta). Seen at Cocha Camungo and Cocha Blanco (oxbow lakes).

Squirrel Cuckoo (Piaya cayana). Common in the Amazon.

Dark-billed Cuckoo (Coccyzus melacoryphus). Seen at Huacarpay lakes.

OWLS

Rufescent Screech-Owl (Megascops ingens). Seen at Cock-of-the-Rock Lodge.

Crested Owl (Lophostrix cristata). Heard only.

Spectacled Owl (Pulsatrix perspicillata). Heard only.

Black-banded Owl (Ciccaba huhula). Seen at Villa Carmen.

Yungas Pygmy-Owl (Glaucidium bolivianum) Seen along the Manu Road. Range Restricted.

Amazonian Pygmy-Owl (Glaucidium hardyi). Heard only.

Ferruginous Pygmy-Owl (Glaucidium brasilianum). Seen around La Cachuela (fish farm) in Puerto Maldonado.

Burrowing Owl (Athene cunicularia). Seen along the Madre de Dios river.

NIGHTJARS

Sand-colored Nighthawk (Chordeiles rupestris). Common. Seen along the Madre de Dios river.

Band-winged Nightjar (Caprimulgus longirostris). Seen near Wayqecha.

Common Nighthawk (Chordeiles minor). Seen at Manu Wildlife Center. A Boreal migrant.

Common Pauraque (Nyctidromus albicollis). Common in the Amazon.

Swallow-tailed Nightjar (Uropsalis segmentata). Seen at Pillahuata area (Manu road).

Lyre-tailed Nightjar (Uropsalis lyra). Seen near Manú Cloud Forest Lodge (1500-1700m) (Manu road).

Ladder-tailed Nightjar (Hydropsalis climacocerca). Seen at Cocha Camungo (oxbow lake).

Ocellated Poorwill (Nyctiphrynus ocellatus). Heard only.

POTOOS

Great Potoo (Nyctibius grandis). Heard only.

Long-tailed Potoo (Nyctibius aethereus). Seen at Manu Wildlife Center.

Common Potoo (Nyctibius griseus). Heard only.

Andean Potoo (Nyctibius maculosus). Seen at Rocotal area.

SWIFTS

Chestnut-collared Swift (Streptoprocne rutila). Seen at Amazonia Lodge.

White-collared Swift (Streptoprocne zonaris). A common species from the foothills down to the Amazon.

Short-tailed Swift (Chaetura brachyura). Seen at Patria (600-900m) (Manu road).

White-tipped Swift (Aeronautes montivagus). Seen along the road between Cusco and Paucartambo.

Lesser Swallow-tailed Swift (Panyptila cayennensis). Seen at Patria (600-900m) (Manu road).

Fork-tailed Palm-Swift (Tachornis squamata). Common in the Amazon. Mainly seen along the Madre de Dios river.

HUMMINGBIRDS

Rufous-breasted Hermit (Glaucis hirsuta). Seen at Manu Wildlife Center.

White-necked Jacobin (Florisuga mellivora). Seen at Amazonia Lodge and Cocha Blanco (oxbow lake).

Buff-tailed Sicklebill (Eutoxeres condamini). Seen from the Tanager Corner to Thousand-meter Bridge (1100-1300m) (Manu road).

White-bearded Hermit (Phaethornis hispidus). Seen at Amazonia Lodge and Manu Wildlife Center.

Koepcke's Hermit (Phaethornis koepckeae). Seen at Amazonia Lodge. ENDEMIC.

Green Hermit (Phaethornis guy). Seen at Cock-of-the-Rock Lodge and Quita Calzones bridge.

Great-billed Hermit (Phaethornis malaris). Seen at Amazonia Lodge.

Reddish Hermit (Phaethornis ruber). Seen at Pastora road in Puerto Maldonado.

White-browed Hermit (Phaethornis stuarti). Seen at Villa Carmen. Named for A.M. Stuart (fl 1897) a collector in Bolivia. Range Restricted.

Green-fronted Lancebill (Doryfera ludovicae). Seen at Cock-of-the-rock Lodge.

Geoffroy's Daggerbill (Schistes geoffroyi). Seen at Cock-of-the-Rock Lodge. Named for Etienne Geoffroy Saint-Hillaire (1772-1884) French zoologist.

Sparkling Violetear (Colibri coruscans). Seen at Cock-of-the-Rock Lodge.

Lesser Violetear (Colobri cyanotus). Seen in Rocotal area.

Black-eared Fairy (Heliothryx auritus). Seen at Amazonia Lodge.

Black-throated Mango (Anthracothorax nigricollis). Seen at La Cachuela.

Amethyst-throated Sunangel (Heliangelus amethysticollis). Fairly common in humid montane forest.

Wire-crested Thorntail (Discosura popelairii). Seen at Cock-of-the-Rock Lodge. Named for Jean Baptiste Baron Popelaire de Terloo Belgian naturalist and collector in Peru in the first half of the 19th century.

Rufous-crested Coquette (Lophornis delattrei). Seen at Amazonia Lodge. Named after French naturalist/collector Henri de Lattre (1838).

Festive Coquette (Lophornis chalybeus). Seen at Manu Wildlife Center.

Peruvian Piedtail (Phlogophilus harterti). Seen at Cock-of-the-Rock Lodge. ENDEMIC.

Speckled Hummingbird (Adelomyia melanogenys). Seen at Cock-of-the-Rock Lodge.

Green-tailed Trainbearer (Lesbia nuna). Seen at Huacarpay lakes. Greek: Lesbia – a woman of Lesbos. Nuna – from Nouna-Koali a graceful Indian virgin in Jean F. Denis’s (1829) novel.

Rufous-capped Thornbill (Chalcostigma ruficeps). Seen near Pillahuata.

(Eastern) Bearded Mountaineer (Oreonympha nobilis). Seen near Huancarani. Sometimes considered a separate species. ENDEMIC.

Long-tailed Sylph (Aglaiocercus kingii). Seen at Buenos Aires area (Manu road) and Cock-of-the-Rock Lodge. Named for Rear-Admiral Philip Parker King (1791-1856) British marine surveyor and collector in tropical America.

Bronzy Inca (Coeligena coeligena). Seen in Rocotal area and at Cock-of-the-Rock Lodge.

Scaled Metaltail (Metallura aeneocauda). Seen at Acjanaco pass to Wayquecha (Manu road). Range Restricted.

Buff-thighed Puffleg (Haplophaedia assimilis). Seen at Pillahuata Tunnels (2200-2500m) (Manu road). Range Restricted.

Shining Sunbeam (Aglaeactis cupripennis). Seen at Acjanaco pass to Wayquecha (Manu road).

Goulds (Collared Inca) (Coeligena (torquata) inca. Seen at Rocotal area. Range Restricted.

Great Sapphirewing (Pterophanes cyanopterus). Seen at Acjanaco pass.

Chestnut-breasted Coronet (Boissonneaua matthewsii). Seen near Pillahuata.

Booted (Anna’s) Racket-tail (Ocreatus underwoodii). Seen at Cock-of-the-Rock Lodge. The Racket tailed Puffleg was unknown in life but specimens existed in various London cabinets, whence a drawing was sent in 1832 by Mr. Underwood on behalf of Charles Stokes, a London stockbroker and collector. ENDEMIC.

Gould's Jewelfront (Heliodoxa aurescens). Seen at Amazonia Lodge.

Violet-fronted Brilliant (Heliodoxa leadbeateri). Seen at Cock-of-the-Rock Lodge.

Giant Hummingbird (Patagona gigas). Seen at Huacarpay. One of the world's largest Hummingbirds after the Sword-billed Hummingbird if talking about measurement from tail tip to bill tip.

Amethyst Woodstar (Calliphlox amethystina). Seen at Amazonia Lodge.

Blue-tailed Emerald (Chlorostilbon mellisugus). Seen at Amazonia Lodge.

Violet-headed Hummingbird (Klais guimeti). Seen at Amazonia Lodge.

Gray-breasted Sabrewing (Campylopterus largipennis). Seen at Amazonia Lodge.

Fork-tailed Woodnymph (Thalurania furcata). Seen at Cock-of-the-Rock Lodge, Amazonia Lodge, and Manu Wildlife Center.

Many-spotted Hummingbird (Taphrospilus hypostictus). Seen at Cock-of-the-Rock Lodge.

White-bellied Hummingbird (Amazilia chionogaster). Seen at Paucartambo.

Sapphire-spangled Emerald (Amazilia lactea). Seen at Amazonia Lodge.

Golden-tailed Sapphire (Chrysuronia oenone). Seen at Amazonia Lodge.

White-chinned Sapphire (Hylocharis cyanus). Seen at Manu Wildlife Center.

QUETZALS AND TROGONS

Golden-headed Quetzal (Pharomachrus auriceps). Seen at Pillahuata Tunnels (2200-2500m) (Manu road), Rocotal area (Manu road) and Mirador (1700-1800m) (Manu road). From the Greek Pharo = Mantle or Cloak. Machrus = long.

Pavonine Quetzal (Pharomacrus pavoninus). Heard only.

Black-tailed Trogon (Trogon melanurus). Common. Mainly seen at Amazonia Lodge.

Amazonian Trogon (Trogon ramonianus). Seen nearby Cocha Camungo (oxbow lake). The subspecies ramonianus and caligatus were formerly (e.g., Cory 1919, Pinto 1937) considered separate species from Trogon violaceus (Violaceous Trogon) but Peters (1945) considered them all conspecific. Ridgely & Greenfield (2001) considered caligatus (Gartered Trogon) of Middle America and northwestern South America to be a separate species from Trogon violaceus, and this was followed by Hilty (2003); Genetic data (DaCosta & Klicka 2008) indicate that caligatus is basal to a group that includes Amazonian T. violaceus, T. curucui, and T. surrucura (and that Amazonian violaceus may be paraphyletic with respect to the latter two species). SACC proposal was passed to recognize caligatus as a species. SACC proposal passed to recognize ramonianus as a separate species from T. violaceus.

Blue-crowned Trogon (Trogon curucui). Seen at Amazonia Lodge and Cocha Camungo (oxbow lake).

Collared Trogon (Trogon collaris). Seen at Amazonia Lodge, Blanquillo Claylick, and Manu Wildlife Center.

Masked Trogon (Trogon personatus). Seen at Pillahuata Tunnels (2200-2500m) (Manu road), Cock-of-the-Rock Lodge, and Quita Calzones bridge.

MOTMOTS

Amazonian Motmot (Momotus momota). Seen at Manu Wildlife Center.

Broad-billed Motmot (Electron platyrhynchum). Heard only.

KINGFISHERS

Ringed Kingfisher (Megaceryle torquata). Seen along the Madre de Dios river and Cocha Camungo (oxbow lake).

Amazon Kingfisher (Chloroceryle amazona). Common in the Amazon. Mainly seen along the Madre de Dios river.

Green Kingfisher (Chloroceryle americana). Seen at Blanquillo Claylick.

American Pygmy Kingfisher (Chloroceryle aenea). Seen at Cocha Camungo.

PUFFBIRDS

Chestnut-capped Puffbird (Bucco macrodactylus). Seen at Amazonia lodge and Villa Carmen.

Black-streaked Puffbird (Malacoptila fulvogularis). Heard only.

Rufous-capped Nunlet (Nonnula ruficapilla). Heard only.

Black-fronted Nunbird (Monasa nigrifrons). Common in the Amazon. Monasa is Greek for solitary or a monk a reference to the plain plumage and quiet behavior of the Nunbirds.

White-fronted Nunbird (Monasa morphoeus). Seen at Manu Wildlife Center. Morpheous was the son of sleep and god of dreams a referral to the lethargic behavior of this Nunbird.

Swallow-winged Puffbird (Chelidoptera tenebrosa). Common in the Amazon. Mainly seen along the Madre de Dios river.

JACAMARS

Purus Jacamar (Galbalcyrhynchus purusianus). Seen at Cocha Camungo and Cocha Blanco (oxbow lakes). Named for the Purus River that runs from eastern Peru into Brazil.

White-throated Jacamar (Brachygalba albogularis). Seen at La Pastora.

Bluish-fronted Jacamar (Galbula cyanescens). Common in the Amazon. Mainly seen at Manu Wildlife Center.

Paradise Jacamar (Galbula dea). Seen at Manu Wildlife Center.

BARBETS

Gilded Barbet (Capito auratus). Heard only.

Lemon-throated Barbet (Eubucco richardsoni). Seen at Amazonia Lodge and Manu Wildlife Center.

Versicolored Barbet (Eubucco versicolor). Seen at Cock-of-the-Rock Lodge.

TOUCANS

Chestnut-tipped Toucanet (Aulacorhynchus derbianus). Seen from the Tanager Corner to Thousand-meter Bridge (1100-1300m) (Manu road).

Gray-breasted Mountain-Toucan (Andigena hypoglauca). Seen from Acjanaco pass to Wayquecha (Manu road).

Golden-collared Toucanet (Selenidera reinwardtii). Seen at Manu Wildlife Center.

Chestnut-eared Aracari (Pteroglossus castanotis). Seen from Thousand-meter Bridge to Chontachaca (900-1100m) (Manu road) and along the Madre de Dios river. In Greek Pteroglossus means “feathertongued” a reference to the slim feather-like tongues of toucans and aracaris.

Curl-crested Aracari (Pteroglossus beauharnaesii). Seen at Manu Wildlife Center.

White-throated Toucan (Ramphastos tucanus). Common in the Amazon. Mainly seen along the Madre de Dios river.

Channel-billed Toucan (Ramphastos vitellinus). Seen at Blanquillo Claylick.

WOODPECKERS

Rufous-breasted Piculet (Picumnus rufiventris). Seen in Antthrush Trail (Manu Wildlife Center).

Fine-barred Piculet (Picumnus subtilis). Heard only.

Yellow-tufted Woodpecker (Melanerpes cruentatus). Common in the Amazon.

Little Woodpecker (Veniliornis passerinus). Seen at Amazonia Lodge and around La Cachuela (fish farm) in Puerto Maldonado.

Red-stained Woodpecker (Veniliornis affinis). Seen from the Tanager Corner to Thousand-meter Bridge (1100-1300m) (Manu road).

Golden-olive Woodpecker (Colaptes rubiginosus). Seen at Cock-of-the-Rock Lodge.

Crimson-mantled Woodpecker (Colaptes rivolii). Seen at Wayqecha Biological Station.

Andean Flicker (Colaptes rupicola). Seen at Huacarpay lakes.

Lineated Woodpecker (Dryocopus lineatus). Seen along the Madre de Dios river, Blanquillo Claylick, and Manu Wildlife Center.

Red-necked Woodpecker (Campephilus rubricollis). Seen at Manu Wildlife Center.

Crimson-crested Woodpecker (Campephilus melanoleucos). Seen along the Madre de Dios river.

FALCONS AND CARACARAS

Laughing Falcon (Herpetotheres cachinnans). Seen at the Blanquillo Claylick.

Barred Forest-Falcon (Micrastur ruficollis). Seen at Manu Wildlife Center.

Collared Forest-Falcon (Micrastur semitorquatus). Seen at Manu Wildlife Center.

Buckley's Forest-Falcon (Micrastur buckleyi). Heard only.

Black Caracara (Daptrius ater). Common in the Amazon. Mainly seen along the Madre de Dios river.

Red-throated Caracara (Ibycter americanus). Seen at Amazonia Lodge, Blanquillo Claylick, Manu Wildlife Centern and Pastora road in Puerto Maldonado.

Mountain Caracara (Phalcoboenus megalopterus). Seen from Acjanaco pass to Wayquecha (Manu road).

Southern Caracara (Caracara plancus). Seen along the Madre de Dios river.

American Kestrel (Falco sparverius). Common in the Andes.

Bat Falcon (Falco rufigularis). Seen along the Madre de Dios river.

Orange-breasted Falcon (Falco deiroleucus). Seen on the Manu Road.

NEW WORLD PARROTS

Scarlet-shouldered Parrotlet (Touit huetii). Seen at Cocha Camungo.

Barred Parakeet (Bolborhynchus lineola). Seen in Pillahuata.

Amazonian Parrotlet (Nannopsittaca dachilleae). Seen at the Blanquillo Claylick. Range Restricted.

Tui Parakeet (Brotogeris sanctithomae). Seen at Blanquillo Claylick.

Cobalt-winged Parakeet (Brotogeris cyanoptera). Common in the Amazon. Mainly seen along the Madre de Dios river.

Blue-headed Parrot (Pionus menstruus). Seen at Blanquillo Claylick.

Speckle-faced Parrot (Pionus tumultuosus). Seen in Pillahuata.

Yellow-crowned Parrot (Amazona ochrocephala). Seen at Blanquillo Claylick.

Mealy Parrot (Amazona farinosa). Seen at Blanquillo Claylick. Farinosa is Latin for “Sprinkled with Flour” referring to the “dusted” appearance of this Amazon.

Scaly-naped Parrot (Amazona mercenarius). Seen at Rocotal area (Manu road).

Dusky-billed Parrotlet (Forpus modestus). Seen at Manu Wildlife Center.

White-bellied Parrot (Pionites leucogaster). Heard only.

Rose-fronted Parakeet (Pyrrhura roseifrons). Seen at Manu Wildlife Center.

Dusky-headed Parakeet (Aratinga weddellii). Common in the Amazon. Mainly seen along the Madre de Dios river.

Red-bellied Macaw (Orthopsittaca manilatus). Seen at Pastora road in Puerto Maldonado.

Blue-headed Macaw (Primolius couloni). Heard only, at Amazonia Lodge.

Blue-and-yellow Macaw (Ara ararauna). Common in the Amazon. Mainly seen along the Madre de Dios river.

Military Macaw (Ara militaris). Seen at Amazonia Lodge.

Scarlet Macaw (Ara macao). Common in the Amazon. Mainly seen along the Madre de Dios river.

Red-and-green Macaw (Ara chloropterus). Common in the Amazon. Mainly seen along the Madre de Dios river and Blanquillo Claylick.

Chestnut-fronted Macaw (Ara severus). Common. Seen from Thousand-meter Bridge to Chontachaca (900-1100m) (Manu road), Quita Calzones bridge, Amazonia Lodge, Manu Wildlife Center, Blanquillo Claylick and along the Madre de Dios river.

White-eyed Parakeet (Psittacara leucophthalmus). Seen from Thousand-meter Bridge to Chontachaca (900-1100m) (Manu road), Quita Calzones bridge, Amazonia Lodge and Manu Wildlife Center.

ANTBIRDS

Chestnut-shouldered Antwren (Euchrepomis humeralis). Seen nearby Cocha Camungo (oxbow lake).

Yellow-rumped Antwren (Euchrepomis sharpei). Seen at Cock-of-the-Rock Lodge. One of the star birds of Manu. Range Restricted.

Bamboo Antshrike (Cymbilaimus sanctaemariae). Seen at Amazonia Lodge.

Great Antshrike (Taraba major). Seen at Pastora road in Puerto Maldonado.

Barred Antshrike (Thamnophilus doliatus). Seen in Puerto Maldonado.

Chestnut-backed Antshrike (Thamnophilus palliatus). Seen at Cock-of-the-Rock Lodge.

Plain-winged Antshrike (Thamnophilus schistaceus). Seen at Amazonia Lodge.

Dusky-throated Antshrike (Thamnomanes ardesiacus). Seen at Manu Wildlife Center.

Bluish-slate Antshrike (Thamnomanes schistogynus). Seen at Amazonia Lodge.

Plain-throated Antwren (Isleria hauxwelli). Seen at Manu Wildlife Center.

Spot-winged Antshrike (Pygiptila stellaris). Heard only.

White-eyed Antwren (Epinecrophylla leucophthalma). Seen at Manu Wildlife Center.

Ornate Antwren (Epinecrophylla ornata). Seen at Amazonia Lodge.

Pygmy Antwren (Myrmotherula brachyura). Seen at Amazonia Lodge.

Sclater's Antwren (Myrmotherula sclateri). Heard only.

Amazonian Streaked-Antwren (Myrmotherula multostriata). Seen at Cocha Blanco.

Stripe-chested Antwren (Myrmotherula longicauda). Seen at Cock-of-the-Rock Lodge.

White-flanked Antwren (Myrmotherula axillaris). Seen at Manu Wildlife Center. Willis (1984), Ridgely & Tudor (1994), Hilty (2003), and Zimmer & Isler (2003) noted that vocal differences among several subspecies of Myrmotherula axillaris suggest that more than one species is involved.

Slaty Antwren (Myrmotherula schisticolor). Seen at Cock-of-the-Rock Lodge.

Long-winged Antwren (Myrmotherula longipennis). Seen at Manu Wildlife Center. One of the common ones in the understory mixed-species flocks.

Gray Antwren (Myrmotherula menetriesii). Seen at Manu Wildlife Center.

Dot-winged Antwren (Microrhopias quixensis). Seen at Amazonia Lodge.

Riparian Antbird (Cercomacroides nigrescens). Heard only.

Black Antbird (Cercomacroides serva). Heard only.

Manu Antbird (Cercomacra manu). Seen in Antthrush Trail (Manu Wildlife Center).

Gray Antbird (Cercomacra cinerascens). Heard only.

White-backed Fire-eye (Pyriglena leuconota). Heard only.

White-browed Antbird (Myrmoborus leucophrys). Seen nearby Cocha Camungo (oxbow lake).

White-lined Antbird (Percnostola lophotes). Seen at Antthrush Trail (Manu Wildlife Center).

Yellow-breasted Warbling-Antbird (Hypocnemis subflava). Seen at Villa Carmen Biological Station.

Peruvian Warbling-Antbird (Hypocnemis peruviana). Seen at Manu Wildlife Center.

Plumbeous Antbird (Myrmelastes hyperythrus). Heard only, at Manu Wildlife Center and Cocha Camungo (oxbow lake).

Chestnut-tailed Antbird (Sciaphylax hemimelaena). Common in the Amazon.

Goeldi's Antbird (Akletos goeldii). Seen in Antthrush Trail (Manu Wildlife Center). Favors Bamboo but is not restricted to it. Named for Emil August Goeldi, German naturalist resident in Brazil and author of “Aves do Brasil” 1894.

Black-throated Antbird (Myrmophylax atrothorax). Seen at Amazonia Lodge.

White-throated Antbird (Oneillornis salvini). Heard only.

Spot-backed Antbird (Hylophylax naevius). Heard only.

Black-spotted Bare-eye (Phlegopsis nigromaculata). Heard only.

GNATEATERS

Slaty Gnateater (Conopophaga ardesiaca). Seen at Cock-of-the-Rock Lodge.

ANTPITTAS

White-throated Antpitta (Grallaria albigula). Heard only.

Red-and-white Antpitta (Grallaria erythroleuca). Seen at Wayqecha Biological Station.

Amazonian Antpitta (Hylopezus berlepschi). Seen at Villa Carmen Biological Station.

Thrush-like Antpitta (Myrmothera campanisona). Heard only.

TAPACULOS

Rusty-belted Tapaculo (Liosceles thoracicus). Seen at Manu Wildlife Center.

Trilling Tapaculo (Scytalopus parvirostris). Heard only.

White-crowned Tapaculo (Scytalopus atratus). Heard only.

ANTTHRUSHES

Black-faced Antthrush (Formicarius analis). Seen at the Antthrush trail.

Rufous-breasted Antthrush (Formicarius rufipectus). Heard only.

Striated Antthrush (Chamaeza nobilis). Seen at Manu Wildlife Center.

OVENBIRDS

Plain-brown Woodcreeper (Dendrocincla fuliginosa). Seen at Manu Wildlife Center.

Wedge-billed Woodcreeper (Glyphorynchus spirurus). Seen at Manu Wildlife Center.

Cinnamon-throated Woodcreeper (Dendrexetastes rufigula). Seen at Amazonia Lodge and Manu Wildlife Center.

Long-billed Woodcreeper (Nasica longirostris). Seen at Manu Wildlife Center.

Strong-billed Woodcreeper (Xiphocolaptes promeropirhynchus). Seen at Pillahuata Tunnels (2200-2500m) (Manu road).

Elegant Woodcreeper (Xiphorhynchus elegans). Heard only.

Buff-throated Woodcreeper (Xiphorhynchus guttatus). Seen at Manu Wildlife Center. The most common Lowland rainforest Woodcreeper; some authorities consider Buff-throated Woodcreeper of SE Brazil as distinct but some authorities consider the reason for splitting weak.

Olive-backed Woodcreeper (Xiphorhynchus triangularis). Seen at Cock-of-the-Rock Lodge and Mirador (1700-1800m) (Manu road).

Straight-billed Woodcreeper (Dendroplex picus). Seen at Pastora road in Puerto Maldonado.

Red-billed Scythebill (Campylorhamphus trochilirostris). Seen at the Antthrush trail.

Montane Woodcreeper (Lepidocolaptes lacrymiger). Seen at Buenos Aires area (Manu road).

Inambari Woodcreeper (Lepidocolaptes fatimalimae). Seen at Manu Wildlife Center.

Streaked Xenops (Xenops rutilans). Seen at Belmond Sanctuary Lodge.

Pale-legged Hornero (Furnarius leucopus). Common in the Amazon.

Wren-like Rushbird (Phleocryptes melanops). Seen at Huacarpay lakes.

Dusky-cheeked Foliage-gleaner (Anabazenops dorsalis). Seen at Amazonia Lodge.

Buff-fronted Foliage-gleaner (Philydor rufum). Seen at Amazonia Lodge.

Montane Foliage-gleaner (Anabacerthia striaticollis). Seen at Cock-of-the-Rock Lodge and Mirador (1700-1800m) (Manu road).

Rufous-tailed Foliage-gleaner (Anabacerthia ruficaudata). Seen at Manu Wildlife Center.

Chestnut-winged Hookbill (Ancistrops strigilatus). Seen at Manu Wildlife Center.

Chestnut-crowned Foliage-gleaner (Automolus rufipileatus). Heard only, at Amazonia Lodge.

Spotted Barbtail (Premnoplex brunnescens). Seen at Cock-of-the-Rock Lodge.

Pearled Treerunner (Margarornis squamiger). Seen from Acjanaco pass to Wayquecha (Manu road).

Streak-fronted Thornbird (Phacellodomus striaticeps). Seen at Huacarpay Lakes.

Puna Thistletail (Asthenes helleri). Seen near Acjanaco. Range Restricted.

Rusty-fronted Canastero (Asthenes ottonis). Seen at Huacarpay Lakes. ENDEMIC.

Orange-fronted Plushcrown (Metopothrix aurantiacus). Seen at Villa Carmen.

Plain Softtail (Thripophaga fusciceps). Seen at Amazonia Lodge.

Creamy-crested Spinetail (Cranioleuca albicapilla). Seen at Acjanaco pass. ENDEMIC.

Ash-browed Spinetail (Cranioleuca curtata). Seen at Cock-of-the-Rock Lodge. Range Restricted.

Plain-crowned Spinetail (Synallaxis gujanensis). Seen at Pastora road in Puerto Maldonado.

Dark-breasted Spinetail (Synallaxis albigularis). Seen along the Madre de Dios River.

Azara's Spinetail (Synallaxis azarae). Seen at Pillahuata.

Cabanis's Spinetail (Synallaxis cabanisi). Heard only.

Plain-crowned Spinetail (Synallaxis gujanensis). Heard only.

TYRANT FLYCATCHERS

White-banded Tyrannulet (Mecocerculus stictopterus). Seen at Pillahuata Tunnels (2200-2500m) (Manu road).

Torrent Tyrannulet (Serpophaga cinerea). Seen in Paucartambo.

White-throated Tyrannulet (Mecocerculus leucophrys). Seen from Acjanaco pass to Wayquecha (Manu road) and Buenos Aires area (Manu road).

Mouse-colored Tyrannulet (Phaeomyias murina). Seen at Cocha Camungo.

Yellow-crowned Tyrannulet (Tyrannulus elatus). Seen at Amazonia Lodge.

White-crested Elaenia (Elaenia albiceps). Seen at Huacarpay Lakes.

Mottle-backed Elaenia (Elaenia gigas). Seen at Amazonia Lodge and nearby Blanquillo Claylick.

Sierran Elaenia (Elaenia pallatangae). Fairly common in humid montane forest.

Streak-necked Flycatcher (Mionectes striaticollis). Seen in humid montane forest.

Ochre-bellied Flycatcher (Mionectes oleagineus). Seen at Manu Wildlife Center.

Slaty-capped Flycatcher (Leptopogon superciliaris). Fairly common. Mainly seen at Cock-of-the-Rock Lodge.

Amazonian Scrub-Flycatcher (Sublegatus obscurior). Seen at Cocha Camungo.

Many-colored Rush-Tyrant (Tachuris rubrigastra). Seen at Huacarpay lakes.

Mottle-cheeked Tyrannulet (Phylloscartes ventralis). Seen at Mirador (1700-1800m) (Manu road).

Cinnamon-faced Tyrannulet (Phylloscartes parkeri). Seen at Quita Calzones bridge. One of the several birds named for Theodore A Parker the III –pioneer US ornithologist tragically killed in plane crash in souther Ecuador in 1993.

Ornate Flycatcher (Myiotriccus ornatus). Seen at Quita Calzones bridge.

Ringed Antpipit (Corythopis torquatus). Seen at Manu Wildlife Center. An atypical ground-dwelling flycatcher.

Bolivian Tyrannulet (Zimmerius bolivianus). Fairly common on east slope of southern Andes.

Variegated Bristle-Tyrant (Phylloscartes poecilotis). Seen in Rocotal.

Streak-necked Flycatcher (Mionectes striaticollis). Widespread Andean flycatcher.

Olive-striped Flycatcher (Mionectes olivaceus). Seen at San Pedro.

McConnell's Flycatcher (Mionectes macconnelli). Seen at Amazonia Lodge.

Inca Flycatcher (Leptopogon taczanowskii). Seen at Rocotal. ENDEMIC.

Sepia-capped Flycatcher (Leptopogon amaurocephalus). Seen at Amazonia Lodge.

Short-tailed Pygmy-Tyrant (Myiornis ecaudatus). Seen at Amazonia Lodge.

Scale-crested Pygmy-Tyrant (Lophotriccus pileatus). Seen at Cock-of-the-Rock Lodge.

Johannes's Tody-Tyrant (Hemitriccus iohannis). Seen in Puerto Maldonado. Range Restricted.

White-bellied Tody-Tyrant (Hemitriccus griseipectus). Seen at Manu Wildlife Center.

Black-throated Tody-Tyrant (Hemitriccus granadensis). Seen in Pillahuata.

Rusty-fronted Tody-Flycatcher (Poecilotriccus latirostre). Heard only.

Black-backed Tody-Flycatcher (Poecilotriccus pulchellus). Heard only. ENDEMIC.

Spotted Tody-Flycatcher (Todirostrum maculatum). Seen at Blanquillo Claylick.

Fulvous-breasted Flatbill (Rhychocyclus fulvipectus). Seen at Cock-of-the-Rock Lodge.

Gray-crowned Flycatcher (Tolmomyias poliocephalus). Seen in Puerto Maldonado.

Yellow-breasted Flycatcher (Tolmomyias viridiceps). Seen at Amazonia Lodge.

Yellow-browed Tody-Flycatcher (Todirostrum chrysocrotaphum). Seen at Manu Wildlife Center.

Golden-crowned Spadebill (Platyrinchus coronatus). Heard only, at Manu Wildlife Center.

Ruddy-tailed Flycatcher (Terenotriccus erythrurus). Seen nearby Cocha Camungo (oxbow lake).

Cinnamon Flycatcher (Pyrrhomyias cinnamomeus). Common in the foothills.

Bran-colored Flycatcher (Myiophobus fasciatus). Seen at Blanquillo Claylick.

Fuscous Flycatcher (Cnemotriccus fuscatus fuscaatior). Seen in Puerto Maldonado. Pronounced differences in vocalizations, habitat, and nest construction indicate that widespread C. f. bimaculatus should be treated as a separate species from nominate Cnemotriccus f. fuscatus (Belton 1984). Hilty (2003) also suggested that C. f. duidae should be treated as a separate species based on voice. In Peru, the taxaduidaeandfuscatiordiffer in voice and occur in close proximity are separated by habitat differences (Alvarez et al. 2012, Harvey et al. 2014, Moncrieff et al. 2019). Clearly, two or more species are currently included within currently defined Cnemotriccus fuscatus, but formal recognition awaits a detailed study that treats all taxa in the complex.

Western Wood-Pewee (Contopus sordidulus). Seen at Cock-of-the-Rock Lodge.

Black Phoebe (Sayornis nigricans). Common. The change from the darker-winged nominate nigricans to the white-winged latirostris is a north-south cline, with larger amounts of white gradually appearing further south.

Vermilion Flycatcher (Pyrocephalus rubinus). Seen in Puerto Maldonado. Austral migrant. In a recent paper – Carmi el al 2016 it strongly suggests several taxonomic changes to the genus Pyrocephalus, including elevating three currently recognized subspecies to full species status: the austral migrant South American subspecies rubinus, and the two Galápagos subspecies dubius and nanus. As such, they propose a revised taxonom.

Drab Water Tyrant (Ochthornis littoralis). Common in the Amazon. Mainly seen along the Madre de Dios river.

Rufous-bellied Bush-Tyrant (Myiotheretes fuscorufus). Seen at Pillahuata Tunnels (2200-2500m) (Manu road).

Slaty-backed Chat-Tyrant (Ochthoeca cinnamomeiventris). Heard only.

Rufous-breasted Chat-Tyrant (Ochthoeca rufipectoralis). Seen in Pillahuata.

Brown-backed Chat-Tyrant (Ochthoeca fumicolor). Seen from Acjanaco pass to Wayquecha (Manu road).

Long-tailed Tyrant (Colonia colonus). Seen at Patria (600-900m) (Manu road), Amazonia Lodge and along the Madre de Dios river.

Rusty-margined Flycatcher (Myiozetetes cayanensis). Seen in Puerto Maldonado.

Rufous-tailed Flatbill (Ramphotrigon ruficauda). Seen at Manu Wildlife Center.

White-rumped Sirystes (Sirystes albocinereus). Seen at Manu Wildlife Center.

Grayish Mourner (Rhytipterna simplex). Heard only, at Manu Wildlife Center.

Dusky-capped Flycatcher (Myiarchus tuberculifer). Seen at Cock-of-the-Rock Lodge, Rocotal area (Manu road) and Pillahuata Tunnels (2200- 2500m) (Manu road).

Short-crested Flycatcher (Myiarchus ferox). Seen at Amazonia Lodge, Blanquillo Claylick, Manu Wildlife Center and along the Madre de Dios river.

Lesser Kiskadee (Pitangus lictor). Seen at the Blanquillo Claylick.

Great Kiskadee (Pitangus sulphuratus). Common in the Amazon. Mainly seen in oxbow lakes.

Boat-billed Flycatcher (Megarynchus pitangua). Seen at Manu Wildlife Center.

Social Flycatcher (Myiozetetes similis). Common in the Amazon.

Gray-capped Flycatcher (Myiozetetes granadensis). Seen from Thousand-meter Bridge to Chontachaca (900-1100m) (Manu road), Patria (600-900m) (Manu road) and Amazonia Lodge.

Lemon-browed Flycatcher (Conopias cinchoneti). Seen at Quita Calzones bridge.

Golden-crowned Flycatcher (Myiodynastes chrysocephalus). Seen at San Pedro.

Streaked Flycatcher (Myiodynastes maculatus). Heard only.

Crowned Slaty Flycatcher (Empidonomus aurantioatrocristatus). Seen at Blanquillo Claylick. An Austral migrant and the bird with the longest scientific name in the world!

Tropical Kingbird (Tyrannus melancholicus). Common in the Amazon.

Short-crested Flycatcher (Myiarchus ferox). Most common and widely distributed Amazonian Myiarchus.

Large-headed Flatbill (Ramphotrigon megacephalum). Seen at Antthrush Trail.

Dull-capped Attila (Attila bolivianus). Heard only, at Blanquillo Claylick.

Bright-rumped Attila (Attila spadiceus). Heard only, at Amazonia Lodge and Manu Wildlife Center.

COTINGAS

Barred Fruiteater (Pipreola arcuata). Seen at Pillahuata.

Andean Cock-of-the-Rock (Rupicola peruvianus). Widespread along east slope of Andes and outlying ridges. Emblem of humid montane forests; often proclaimed as Peru's national bird.

Purple-throated Fruitcrow (Querula purpurata). Seen at Manu Wildlife Center.

Amazonian Umbrellabird (Cephalopterus ornatus). Seen at San Pedro.

Screaming Piha (Lipaugus vociferans). Common and widespread. The voice of Amazonia: its far-carrying song is one of the most characteristic sounds of lowland forest.

Bare-necked Fruitcrow (Gymnoderus foetidus). Seen at Manu Wildlife Center.

MANAKINS

Yungas Manakin (Chiroxiphia boliviana). Heard only.

Fiery-capped Manakin (Machaeropterus pyrocephalus). Seen at Amazonia Lodge.

Band-tailed Manakin (Pipra fasciicauda). Fairly common in Amazonia.

Cerulean-capped Manakin (Lepidothrix coeruleocapilla). Seen at Cock-of-the-Rock Lodge. ENDEMIC.

TITYRAS AND BECARDS

Masked Tityra (Tityra semifasciata). Seen at the Blanquillo Claylick.

Black-crowned Tityra (Tityra inquisitor). Seen at the Blanquillo Claylick.

Varzea Schiffornis (Schiffornis major). Seen at Cocha Camungo.

Brown-winged Schiffornis (Schiffornis turdinus). Seen at Manu Wildlife Center.

Barred Becard (Pachyramphus versicolor). Seen at Pillahuata.

VIREOS

Dusky-capped Greenlet (Pachysylvia hypoxantha). Heard only.

Chivi Vireo (Vireo chivi). Seen at Amazonia.

JAYS

White-collared Jay (Cyanolyca viridicyanus). Seen at Buenos Aires area (Manu road) and Pillahuata Tunnels (2200-2500m) (Manu road).

Purplish Jay (Cyanocorax cyanomelas). Seen at Amazonia Lodge and along the Madre de Dios river.

Violaceous Jay (Cyanocorax violaceus). Common in the Amazon.

SWALLOWS

Blue-and-white Swallow (Pygochelidon cyanoleuca). Common.

Brown-bellied Swallow (Orochelidon murina). Seen at Paucartambo (town) and Acjanaco pass to Wayquecha (Manu road).

White-thighed Swallow (Atticora tibialis). Seen at Amazonia Lodge.

White-banded Swallow (Atticora fasciata). Common in the Amazon. Mainly seen along the Madre de Dios river.

Southern Rough-winged Swallow (Stelgidopteryx ruficollis). Fairly common in the Amazon. Mainly seen along the Madre de Dios river.

Gray-breasted Martin (Progne chalybea). Seen at Cocha Blanco (oxbow lake).

Brown-chested Martin (Progne tapera). Uncommon. Seen along the Madre de Dios river.

White-winged Swallow (Tachycineta albiventer). Common in the Amazon. Mainly seen along the Madre de Dios river.

WRENS

Scaly-breasted Wren (Microcerculus marginatus). Heard only.

Gray-mantled Wren (Odontorchilus branickii). Seen at Cock-of-the-Rock Lodge.

House Wren (Troglodytes aedon). Common. Many authors (e.g., Hellmayr 1934, Pinto 1944, Phelps & Phelps 1950a) formerly treated Neotropical mainland populations as a separate species T. musculus; see also Brumfield and Capparella (1996); this treatment was followed by Brewer (2001) and Kroodsma & Brewer (2005). The Falklands population, T. a. cobbi, is treated as a species (Wood 1993), as was done by Brewer (2001), Mazar Barnett & Pearman (2001), Jaramillo (2003), and Kroodsma & Brewer (2005).

Mountain Wren (Troglodytes solstitialis). Seen at Pillahuata Tunnels (2200-2500m) (Manu road).

Sedge Wren (Cistothorus platensis). Seen near Acjanaco Pass.

Thrush-like Wren (Campylorhynchus turdinus). Seen at Manu Wildlife Center.

Moustached Wren (Pheugopedius genibarbis). Seen at Amazonia Lodge.

Buff-breasted Wren (Cantorchilus leucotis). Seen around La Cachuela (fish farm) in Puerto Maldonado.

Fulvous Wren (Cynnycerthia fulva). Seen at Pillahuata. Range Restricted.

Gray-breasted Wood-Wren (Henicorhina leucophrys). Seen at Rocotal.

Chestnut-breasted Wren (Cyphorhinus thoracicus). Heard only.

DIPPERS

White-capped Dipper (Cinclus leucocephalus). Seen at Cock-of-the-Rock Lodge.

DONACOBIUS

Black-capped Donacobius (Donacobius atricapilla). Fairly common. Seen at oxbow lakes.

THRUSHES AND SOLITAIRES

Andean Solitaire (Myadestes ralloides). Seen at Cock-of-the-Rock Lodge.

Black-billed Thrush (Turdus ignobilis). Common in the Amazon.

Chiguanco Thrush (Turdus chiguanco). Common in the Andes.

Great Thrush (Turdus fuscater). Seen at Wayqecha.

White-necked Thrush (Turdus albicollis). Seen at Manu Wildlife Center. Guide only.

NEW WORLD WARBLERS

Tropical Parula (Setophaga pitiayumi). Fairly common in humid montane forest along lower east slope of Andes and on outlying ridges.

Citrine Warbler (Myiothlypis luteoviridis). Seen at Pillahuata.

Pale-legged Warbler (Myiothlypis signatus). Seen at Pillahuata.

Buff-rumped Warbler (Myiothlypis fulvicauda). Seen at Manu Wildlife Center.

Two-banded Warbler (Myiothlypis bivittata). Fairly common in southern Andes, in understory of humid forest and at forest edge.

Russet-crowned Warbler (Myiothlypis coronatus). Seen at Cock-of-the-Rock Lodge.

Three-striped Warbler (Basileuterus tristriatus tristriatus). Seen at San Pedro.

Slate-throated Redstart (Myioborus miniatus). Common in humid montane forest.

Spectacled Redstart (Myioborus melanocephalus). Common in humid montane forest.

TANAGERS AND ALLIES

Red-capped Cardinal (Paroaria gularis). Common in the Amazon.

Magpie Tanager (Cissopis leverianus). Seen at Cocha Blanco (oxbow lake).

Hooded Tanager (Nemosia pileata). Seen near Blanquillo Claylick.

Slaty Tanager (Creurgops dentate). Seen at Pillahuata.

Drab Hemispingus (Pseudospingus xanthophthalmus). Seen at Acjanaco Pass.

Superciliaried Hemispingus (Thlypopsis superciliaris). Seen at Pillahuata.

Rust-and-yellow Tanager (Thlypopsis ruficeps). Fairly common along east slope of Andes in montane shrubbery, at forest edge, and in bamboo thickets; locally also in Polylepis woods.

Black-goggled Tanager (Trichothraupis melanops). Seen at San Pedro.

White-shouldered Tanager (Tachyphonus luctuosus). Fairly common and widespread in Amazonia.

White-winged Shrike-Tanager (Lanio versicolor). Seen at Cocha Camungo.

Silver-beaked Tanager (Ramphocelus carbo). Widespread and common in second growth and forest edge throughout Amazonia.

Masked Crimson Tanager (Ramphocelus nigrogularis). Fairly common in the Amazon. Mainly seen at Amazonia Lodge.

Hooded Mountain-Tanager (Buthraupis montana). Fairly common in humid montane forest along east slope of Andes.

Grass-green Tanager (Chlorornis riefferii). Seen near Wayqecha.

Scarlet-bellied Mountain-Tanager (Anisognathus igniventris). Fairly common in humid montane forest.

Chestnut-bellied Mountain-Tanager (Dubusia castaneoventris). Seen at Rocotal.

Yellow-throated Tanager (Iridosornis analis). Seen at Cock-of-the-Rock Lodge and Mirador (1700-1800m) (Manu road).

Golden-collared Tanager (Iridosornis jelskii). Seen at Acjanaco pass to Wayquecha (Manu road).

Blue-and-yellow Tanager (Pipraeidea bonariensis). Fairly common and widespread in Andes in dry montane scrub, forest edge, and agricultural areas; locally descends to coast in central Peru and and on east slope of Andes.

Orange-eared Tanager (Chlorochrysa calliparaea). Seen at Cock-of-the-Rock Lodge.

Blue-gray Tanager (Thraupis episcopus). Common in the Amazon. Episcopus – a reference to the episcopal blue plumage of this species.

Palm Tanager (Thraupis palmarum). Common in the Amazon.

Blue-capped Tanager (Thraupis cyanocephala). Fairly common in humid montane forest.

Golden-naped Tanager (Chalcothraupis ruficervix). Seen at Pillahuata.

Masked Tanager (Tangara nigrocincta). Seen at Amazonia Lodge.

Spotted Tanager (Ixothraupis punctata). Seen at Cock-of-the-Rock Lodge.

Blue-and-black Tanager (Tangara vassorii). Seen at Rocotal.

Blue-necked Tanager (Tangara cyanicollis). Fairly common in the foothills.

Beryl-spangled Tanager (Tangara nigroviridis). Seen at Rocotal area (Manu road) and Cock-of-the-Rock Lodge.

Paradise Tanager (Tangara chilensis). Common in the Amazon. Not found in Chile as the species name refers.

Bay-headed Tanager (Tangara gyrola). Seen at San Pedro.

Golden-eared Tanager (Tangara chrysotis). Seen at Cock-of-the-Rock Lodge.

Saffron-crowned Tanager (Tangara xanthocephala). Seen at Buenos Aires area (Manu road) and Rocotal area (Manu road). Here the subspecies has an orange not saffron crown.

Green-and-gold Tanager (Tangara schrankii). Seen at Thousand-meter Bridge to Chontachaca (900-1100m) (Manu road), Amazonia Lodge and Manu Wildlife Center.

Golden Tanager (Tangara arthus). Seen at Cock-of-the-Rock Lek (Manu road).

Swallow Tanager (Tersina viridis). Seen at Manu Wildlife Center and Blanquillo Claylick.

Black-faced Dacnis (Dacnis lineata). Seen at Amazonia Lodge and nearby Cocha Camungo (oxbow lake).

Yellow-bellied Dacnis (Dacnis flaviventer). Seen at Amazonia Lodge and Manu Wildlife Center.

Blue Dacnis (Dacnis cayana). Seen at Thousand-meter Bridge to Chontachaca (900-1100m) (Manu road), Amazonia Lodge and Manu Wildlife Center.

Purple Honeycreeper (Cyanerpes caeruleus). Seen nearby Cocha Camungo (oxbow lake).

Green Honeycreeper (Chlorophanes spiza). Seen nearby Cocha Camungo (oxbow lake).

Golden-collared Honeycreeper (Iridophanes pulcherrimus). Seen at Mirador (1700-1800m) (Manu road).

Blue-backed Conebill (Conirostrum sitticolor). Seen at Acjanaco pass to Wayquecha (Manu road).

Capped Conebill (Conirostrum albifrons). Seen at Pillahuata.

Rusty Flowerpiercer (Diglossa sittoides). Seen at Huacarpay lakes.

Cinereous Conebill (Conirostrum cinereum). Common in the Andes.

White-browed Conebill (Conirostrum ferrugineiventre). Seen at Acjanaco Pass.

Moustached Flowerpiercer (Diglossa mystacalis). Seen at Acjanaco Pass.

Black-throated Flowerpiercer (Diglossa brunneiventris). Seen at Acjanaco Pass.

Bluish Flowerpiercer (Diglossa caerulescens). Seen at Mirador (1700-1800m) (Manu road).

Masked Flowerpiercer (Diglossa cyanea). One of the most frequent members of mixed-species flocks in high elevation, humid montane forest of Andes.

Peruvian Sierra-Finch (Geospizopis unicolor). Common and widespread in Andes.

Mourning Sierra-Finch (Rhopospina fruticeti). Seen at Huacarpay Lakes.

Ash-breasted Sierra-Finch (Geospizopsis plebejus). Seen at Huacarpay Lakes.

Chestnut-breasted Mountain-Finch (Poospizopsis caesar). Seen on the road from Cusco city to Paucartambo. ENDEMIC.

Saffron Finch (Sicalis flaveola). Seen in Puerto Maldonado.

Greenish Yellow-Finch (Sicalis olivacens). Seen at Huacarpay Lakes.

Grassland Yellow-Finch (Sicalis luteola). Seen at Huacarpay Lakes.

Blue-black Grassquit (Volatinia jacarina). Seen in Puerto Maldonado.

Chestnut-bellied Seedeater (Sporophila castaneiventris). Seen at Villa Carmen Biological Station.

Buff-throated Saltator (Saltator maximus). Fairly common in the foothills and the Amazon. Common bird below 1300 meters – seen at Cock of the Rock Lodge, Villa Carmen, Amazonia Lodge and Manu Wildlife Center.

Grayish Saltator (Saltator coerulescens). Seen at Blanquillo Claylick.

Golden-billed Saltator (Saltator aurantiirostris). Seen at Huacarpay Lakes.

Chestnut-bellied Seed-Finch (Sporophila angolensis). Seen at Blanquillo Claylick.

Double-collared Seedeater (Sporophila caerulescens). Seen at Blanquillo Claylick, Cocha Camungo (oxbow lake) and Pastora road in Puerto Maldonado.

Band-tailed Seedeater (Catamenia analis). Seen at Huacarpay Lakes.

Bananaquit (Coereba flaveola). Seen at Quita Calzones bridge and Cock-of-the-Rock Lodge. Not common in Manu.

NEW WORLD SPARROWS AND ALLIES

Yellow-throated Chlorospingus (Chlorospingus flavigularis). Fairly common in humid montane forest. Common in mixed-species flocks.

Common Chlorospingus (Chlorospingus flavopectus). Seen at Buenos Aires area (Manu road).

Yellow-browed Sparrow (Ammodramus aurifrons). Common in the Amazon.

Rufous-collared Sparrow (Zonotrichia capensis). A very common bird of the Andes throughout Peru and South America.

Black-faced Brushfinch (Atlapetes melanolaemus). Seen at Pillahuata Tunnels (2200-2500m) (Manu road), Rocotal area (Manu road) and Mirador (1700-1800m) (Manu road). The Atlapetes genus has been completely revamped based on bio-chemical data and work done at Copenhagen Field Museum. Formerly considered part of Rufous-napped Brush-Finch complex A. rufinucha. Atlapetes melanolaemus was formerly (Hellmayr 1938, Paynter 1970a, Meyer de Schauensee 1970, Ridgely & Tudor 1989, Sibley & Monroe 1990) considered a subspecies of A. rufinucha, but see García-Moreno & Fjeldså (1999).

CARDINALS AND ALLIES

Red-crowned Ant-Tanager (Habia rubica). Seen at Manu Wildlife Center.

BLACKBIRDS, ORIOLES, AND OROPENDOLAS

Russet-backed Oropendola (Psarocolius angustifrons). Common in the Amazon.

Dusky-green Oropendola (Psarocolius atrovirens). Seen at Cock-of-the-Rock Lodge.

Crested Oropendola (Psarocolius decumanus). Seen at Thousand-meter Bridge to Chontachaca (900-1100m) (Manu road), Quita Calzones bridge and along the Madre de Dios river.

Olive Oropendola (Psarocolius bifasciatus). Seen along the Madre de Dios river and Manu Wildlife Center.

Solitary Cacique (Cacicus solitarius). Seen at Amazonia Lodge and Pastora road in Puerto Maldonado.

Yellow-rumped Cacique (Cacicus cela). Common in the Amazon.

Red-rumped Cacique (Cacicus haemorrhous). Seen at Cocha Blanco.

Casqued Cacique (Cacicus oseryi). Seen at Amazonia Lodge.

Mountain Cacique (Cacicus chrysonotus). Seen at Pillahuata.

Epaulet Oriole (Icterus cayanensis). Seen in Puerto Maldonado.

Orange-backed Troupial (Icterus croconotus). Seen at Cocha Blanco (oxbow lake).

Giant Cowbird (Molothrus oryzivorus). Common in the Amazon. Mainly seen along the Madre de Dios river. – A brood parasite.

Pale-eyed Blackbird (Agelasticus xanthophthalmus). Seen at Cocha Blanco (oxbow lake).

Yellow-winged Blackbird (Agelasticus thilius). Seen at Huacarpay Lakes.

FINCHES, EUPHONIAS, AND ALLIES

Blue-naped Chlorophonia (Chlorophonia cyanea). Seen at Cock-of-the-rock Lodge.

Golden-bellied Euphonia (Euphonia chrysopasta). Seen at the Blanquillo Claylick.

Purple-throated Euphonia (Euphonia chlorotica). Seen in Puerto Maldonado.

Bronze-green Euphonia (Euphonia mesochrysa). Seen at Cock-of-the-Rock Lodge. Common in mixed flocks.

Orange-bellied Euphonia (Euphonia xanthogaster). Most common, widespread euphonia. Humid montane and lowland forests.

Hooded Siskin (Spinus magellanicus). Seen at Huacarpay Lakes.

Olivaceous Siskin (Spinus olivaceus). Seen at Pillahauta.